Langbahn Team – Weltmeisterschaft

List of soricines

Brown shrew
Southern water shrew (Neomys anomalus)

Soricinae is a subfamily of small mammals in the shrew family Soricidae, which in turn is part of the order Eulipotyphla. A member of this family is called a soricine or a red-toothed shrew. Soricinae is one of three subfamilies in Soricidae, along with the white-toothed shrews of Crocidurinae and the African shrews of Myosoricinae. They are found in Europe, Asia, North America, and northern South America, primarily in forests, shrublands, grasslands, and wetlands, and also in rocky areas and deserts. They range in size from the Eurasian pygmy shrew, at 3 cm (1 in) plus a 3 cm (1 in) tail, to various members of the Chimarrogale genus, at 14 cm (6 in) plus a 13 cm (5 in) tail. Soricines primarily eat insects and other invertebrates, as well as small vertebrates and plants. No soricines have population estimates, but 6 species are categorized as endangered species: the Bornean water shrew, Darién small-eared shrew, Enders's small-eared shrew, Guatemalan broad-clawed shrew, Sierra de Aroa shrew, and Pribilof Island shrew. Additionally, the Nelson's small-eared shrew, San Cristobal shrew, and Sclater's shrew are categorized as critically endangered.

The 158 extant species of Soricinae are divided into 13 genera; 76 of them are in Sorex and 41 in Cryptotis, with the remaining 41 species divided between the other 11 genera. A few extinct prehistoric soricines have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[1]

Conventions

IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (3 species)
 EN Endangered (6 species)
 VU Vulnerable (6 species)
 NT Near threatened (4 species)
 LC Least concern (103 species)
Other categories
 DD Data deficient (34 species)
 NE Not evaluated (1 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the soricine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

The shrew subfamily Soricinae consists of thirteen genera. The majority of the species are contained within Cryptotis, with 41 species, and Sorex, with 76 species. The remaining shrews are split between Chodsigoa with eight species; Chimarrogale with six; Anourosorex, Episoriculus, Neomys, and Notiosorex with four species each; Blarina and Blarinella with three species each, and Megasorex, Nectogale, and Soriculus each with a single species.

Subfamily Soricinae

  • Genus Anourosorex (Asian mole shrews): four species
  • Genus Blarina (short-tailed shrews): three species
  • Genus Blarinella (Asian short-tailed shrews): three species
  • Genus Chimarrogale (Asiatic water shrews): six species
  • Genus Chodsigoa (Asiatic shrews): eight species
  • Genus Cryptotis (small-eared shrews): 41 species
  • Genus Episoriculus (brown-toothed shrews): four species
  • Genus Megasorex (Mexican shrew): one species
  • Genus Nectogale (Elegant water shrew): one species
  • Genus Neomys (water shrews): four species
  • Genus Notiosorex (gray shrews): four species
  • Genus Sorex (long-tailed shrews): 76 species
  • Genus Soriculus (Himalayan shrew): one species

Soricines

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[5]

Genus Anourosorex A. Milne-Edwards, 1872 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Assam mole shrew


A. assamensis
Anderson, 1875
Northeastern India
Map of range
Size: 8–12 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[6]

Habitat: Forest and rocky areas[7]

Diet: Insects and earthworms[6]
 LC 


Unknown Unknown[7]

Chinese mole shrew


A. squamipes
H. Milne-Edwards, 1872
Eastern Asia
Map of range
Size: 8–11 cm (3–4 in), plus tail[8]

Habitat: Forest[9]

Diet: Insects and worms, as well as plants[8]
 LC 


Unknown Unknown[9]

Giant mole shrew


A. schmidi
Petter, 1963
Northeastern India and Bhutan
Map of range
Size: 8–12 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[6]

Habitat: Forest[10]

Diet: Insects and earthworms[6]
 DD 


Unknown Unknown[10]

Taiwanese mole shrew

Gray shrew

A. yamashinai
Kuroda, 1935
Taiwan
Map of range
Size: 8–12 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[6]

Habitat: Forest and grassland[11]

Diet: Insects and earthworms[6]
 LC 


Unknown Unknown[11]

Genus Blarina Gray, 1838 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Elliot's short-tailed shrew

Gray shrew

B. hylophaga
Elliot, 1899
Central United States
Map of range
Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[12]

Habitat: Grassland, shrubland, and forest[13]

Diet: Insects, arthropods, and earthworms, as well as small vertebrates and plants[12]
 LC 


Unknown Population steady[13]

Everglades short-tailed shrew


B. peninsulae
Merriam, 1895
Florida Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[14]

Habitat: Forest, shrubland, grassland, and inland wetlands[14]

Diet: Invertebrates, small vertebrates, and plants[14]
 NE 


Unknown Unknown

Northern short-tailed shrew

Gray shrew

B. brevicauda
(Say, 1823)
Eastern North America
Map of range
Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[15]

Habitat: Inland wetlands, shrubland, and forest[16]

Diet: Invertebrates, small vertebrates, and plants[15]
 LC 


Unknown Population steady[16]

Southern short-tailed shrew

Gray shrew

B. carolinensis
(Bachman, 1837)

Two subspecies
  • B. c. carolinensis
  • B. c. minima
Southeastern United States
Map of range
Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[17]

Habitat: Forest, shrubland, grassland, and inland wetlands[18]

Diet: Invertebrates, as well as plants[17]
 LC 


Unknown Population steady[18]

Genus Blarinella Thomas, 1911 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Asiatic short-tailed shrew


B. quadraticauda
H. Milne-Edwards, 1872
Central China
Map of range
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[19]

Habitat: Forest and shrubland[20]

Diet: Invertebrates[19]
 NT 


Unknown Unknown[20]

Burmese short-tailed shrew


B. wardi
Thomas, 1915
Southern China and Myanmar
Map of range
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[19]

Habitat: Forest[21]

Diet: Invertebrates[19]
 LC 


Unknown Unknown[21]

Indochinese short-tailed shrew


B. griselda
Thomas, 1912
Central and southern China and northern Vietnam
Map of range
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[19]

Habitat: Forest[22]

Diet: Invertebrates[19]
 LC 


Unknown Unknown[22]

Genus Chimarrogale Anderson, 1877 – six species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bornean water shrew


C. phaeura
Thomas, 1898
Northern Borneo in Malaysia
Map of range
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest and inland wetlands[23]

Diet: Invertebrates[23]
 EN 


Unknown Population declining[23]

Chinese water shrew


C. styani
De Winton, 1899
Central China and Myanmar
Map of range
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest, shrubland, and inland wetlands[24]

Diet: Aquatic insects, crustaceans, and small fish[6]
 LC 


Unknown Unknown[24]

Himalayan water shrew


C. himalayica
(Gray, 1842)
Eastern Asia
Map of range
Size: 11–14 cm (4–6 in) long, plus 7–9 cm (3–4 in) tail[25]

Habitat: Forest and inland wetlands[26]

Diet: Insects and spiders[25]
 LC 


Unknown Unknown[26]

Japanese water shrew

Gray shrew

C. platycephalus
Temminck, 1842
Japan
Map of range
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Inland wetlands[27]

Diet: Aquatic insects, crustaceans, and small fish[6]
 LC 


Unknown Population declining[27]

Malayan water shrew


C. hantu
Harrison, 1958
Malaysia
Map of range
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest and inland wetlands[28]

Diet: Aquatic insects, crustaceans, and small fish[6]
 NT 


Unknown Population declining[28]

Sumatran water shrew


C. sumatrana
(Thomas, 1921)
Southern Sumatra in Indonesia
Map of range
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest and inland wetlands[29]

Diet: Aquatic insects, crustaceans, and small fish[29]
 DD 


Unknown Unknown[29]

Genus Chodsigoa Kastchenko, 1907 – eight species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
De Winton's shrew


C. hypsibia
De Winton, 1899
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest and shrubland[31]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[31]

Lamulate shrew


C. lamula
(Thomas, 1912)
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[32]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[32]

Lesser Taiwanese shrew


C. sodalis
(Thomas, 1913)
Taiwan
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[33]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown Unknown[33]

Lowe's shrew


C. parca
(Allen, 1923)

Three subspecies
  • C. p. furva
  • C. p. lowei
  • C. p. parca
Southern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[34]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[34]

Pygmy brown-toothed shrew


C. parva
Allen, 1923
Southern China
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Unknown[35]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown Unknown[35]

Salenski's shrew


C. salenskii
(Kastchenko, 1907)
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Unknown[36]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown Unknown[36]

Smith's shrew


C. smithii
(Thomas, 1911)
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[37]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 NT 


Unknown Population declining[37]

Van Sung's shrew


C. caovansunga
(Lunde, Musser, & Son, 2003)
Northern Vietnam
Map of range
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[38]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown Unknown[38]

Genus Cryptotis Pomel, 1848 – 41 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Big Mexican small-eared shrew


C. magna
(Merriam, 1895)
Southern Mexico
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[40]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 VU 


Unknown Population declining[40]

Blackish small-eared shrew


C. nigrescens
(J. A. Allen, 1895)
Central America
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[41]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[41]

Blind small-eared shrew


C. niausa
Moreno Cárdenas & Albuja, 2014
Northern Ecuador Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest, grassland, and rocky areas[42]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[42]

Celaque broad-clawed shrew


C. celaque
Woodman, 2015
Western Honduras Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[43]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[43]

Central American least shrew

Brown shrew

C. orophila
(Allen, 1895)
Central America
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest, shrubland, and inland wetlands[44]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[44]

Central Mexican broad-clawed shrew


C. alticola
(Merriam, 1895)
Central Mexico
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[45]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[45]

Colombian small-eared shrew


C. colombiana
Woodman & Timm, 1993
Central Colombia
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[46]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[46]

Darién small-eared shrew


C. merus
Goldman, 1912
Border of Colombia and Panama
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[47]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown Population declining[47]

Dinira small-eared shrew


C. dinirensis
Quiroga-Carmona & DoNascimiento, 2016
Northern Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[48]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[48]

Eastern Cordillera small-footed shrew


C. brachyonyx
Woodman, 2003
Central Colombia
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Unknown[49]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[49]

Ecuadorian small-eared shrew


C. equatoris
Thomas, 1912
Ecuador
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and unknown[50]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[50]

Enders's small-eared shrew


C. endersi
Setzer, 1950
Panama
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[51]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown Unknown[51]

Goldman's broad-clawed shrew


C. goldmani
(Merriam, 1895)
Southern Mexico
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[52]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[52]

Goodwin's broad-clawed shrew


C. goodwini
Jackson, 1933
Southern Mexico and Central America
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[53]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[53]

Grizzled Mexican small-eared shrew


C. obscura
(Merriam, 1895)
Eastern Mexico Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[54]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population declining[54]

Guatemalan broad-clawed shrew


C. griseoventris
Jackson, 1933
Southern Mexico and Guatemala
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[55]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown Population declining[55]

Honduran small-eared shrew


C. hondurensis
Woodman & Timm, 1992
Honduras
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[56]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[56]

Mam broad-clawed shrew


C. mam
Woodman, 2010
Guatemala Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[57]

Diet: Earthworms, insects, and plants[57]
 DD 


Unknown Unknown[57]

Medellín small-eared shrew

Black shrew

C. medellinia
Thomas, 1921
Colombia
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[58]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[58]

Merida small-eared shrew


C. meridensis
Thomas, 1898
Venezuela
Map of range
Size: 7–11 cm (3–4 in) long, plus 2–5 cm (1–2 in) tail[59]

Habitat: Forest and shrubland[60]

Diet: Insects and other invertebrates, as well as small vertebrates and eggs[59]
 VU 


Unknown Population steady[60]

Merriam's small-eared shrew


C. merriami
Choate, 1970
Southern Mexico and Central America
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[61]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[61]

Mexican small-eared shrew


C. mexicana
(Coues, 1870)
Southern Mexico
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[62]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population declining[62]

Muscular broad-clawed shrew


C. lacertosus
Woodman, 2010
Guatemala Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[63]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[63]

Nelson's small-eared shrew


C. nelsoni
Merriam, 1895
Eastern Mexico
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[64]

Diet: Insects[64]
 CR 


Unknown Unknown[64]

North American least shrew

Gray shrew

C. parva
(Say, 1823)
Eastern North America
Map of range
Size: 7–10 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[65]

Habitat: Forest, shrubland, grassland, inland wetlands, and rocky areas[66]

Diet: Insects, as well as other invertebrates[65]
 LC 


Unknown Population steady[66]

Oaxacan broad-clawed shrew


C. peregrina
(Merriam, 1895)
Southern Mexico Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[67]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[67]

Omoa broad-clawed shrew


C. mccarthyi
Woodman, 2015
Honduras Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[68]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[68]

Perija small-eared shrew


C. perijensis
Quiroga-Carmona & Woodman, 2015
Border of Colombia and Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[69]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[69]

Peruvian small-eared shrew


C. peruviensis
Vivar, Pacheco, & Valqui, 1997
Northern Peru
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[70]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[70]

Phillips's small-eared shrew


C. phillipsii
(Schaldach, 1966)
Southern Mexico Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[71]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 VU 


Unknown Population declining[71]

Santa Barbara broad-clawed shrew


C. cavatorculus
Woodman, 2015
Honduras Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[72]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[72]

Scaly-footed small-eared shrew


C. squamipes
(Allen, 1916)
Colombia and Ecuador
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[73]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Unknown[73]

Sierra de Aroa shrew


C. aroensis
Quiroga-Carmona & Molinari, 2012
Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[74]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown Unknown[74]

Talamancan small-eared shrew


C. gracilis
Miller, 1911
Costa Rica and Panama
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[75]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Unknown[75]

Tamá small-eared shrew


C. tamensis
Woodman, 2002
Border of Colombia and Venezuela
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[76]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Unknown[76]

Thomas's small-eared shrew


C. thomasi
(Merriam, 1897)
Colombia
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[77]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population steady[77]

Tropical small-eared shrew


C. tropicalis
(Merriam, 1895)
Southern Mexico and Central America
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest, grassland, and inland wetlands[78]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[78]

Venezuelan small-eared shrew


C. venezuelensis
Quiroga-Carmona, 2013
Northern Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[79]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[79]

Wandering small-eared shrew


C. montivaga
(Anthony, 1921)
Ecuador
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[80]

Diet: Insects and other invertebrates[80]
 LC 


Unknown Population steady[80]

Yalijux shrew


C. oreoryctes
Woodman, 2011
Guatemala Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[81]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown Unknown[81]

Yucatan small-eared shrew


C. mayensis
(Merriam, 1901)
Southern Mexico and Central America
Map of range
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[82]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown Population declining[82]

Genus Episoriculus Ellerman & Morrison-Scott, 1966 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Hodgson's brown-toothed shrew


E. caudatus
(Horsfield, 1851)
Southern Asia
Map of range
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest, shrubland, and grassland[83]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[83]

Long-tailed brown-toothed shrew


E. leucops
(Horsfield, 1855)
Southern Asia
Map of range
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest[84]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[84]

Long-tailed mountain shrew


E. macrurus
Blanford, 1888
Southern Asia
Map of range
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest[85]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[85]

Taiwanese brown-toothed shrew


E. fumidus
(Thomas, 1913)
Taiwan
Map of range
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest and shrubland[86]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[86]

Genus Megasorex Hibbard, 1950 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Mexican shrew


M. gigas
(Merriam, 1897)
Southern Mexico
Map of range
Size: 8–9 cm (3–4 in) long, plus 3–5 cm (1–2 in) tail[87]

Habitat: Forest and shrubland[88]

Diet: Invertebrates[87]
 LC 


Unknown Unknown[88]

Genus Nectogale A. Milne-Edwards, 1870 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Elegant water shrew


N. elegans
H. Milne-Edwards, 1870
Central China and southern Asia
Map of range
Size: 9–13 cm (4–5 in) long, plus 8–11 cm (3–4 in) tail[89]

Habitat: Forest and inland wetlands[90]

Diet: Insects, crustaceans, and small fish[89]
 LC 


Unknown Unknown[90]

Genus Neomys Kaup, 1829 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Eurasian water shrew

Gray shrew

N. fodiens
(Pennant, 1771)
Europe and Asia
Map of range
Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Forest, grassland, inland wetlands, intertidal marine, and coastal marine[91]

Diet: Aquatic invertebrates, fish, amphibians, and frogs[92]
 LC 


Unknown Population steady[91]

Iberian water shrew


N. anomalus
A. Cabrera, 1907
Western Europe Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Inland wetlands[93]

Diet: Invertebrates and small vertebrates such as fish and frogs[87]
 LC 


Unknown Population declining[93]

Mediterranean water shrew

Gray shrew

N. milleri
Mottaz, 1907
Europe and western Asia
Map of range
Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Inland wetlands[94]

Diet: Invertebrates and small vertebrates such as fish and frogs[87]
 LC 


Unknown Population declining[94]

Transcaucasian water shrew


N. teres
Miller, 1908
Western Asia
Map of range
Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Forest and inland wetlands[95]

Diet: Invertebrates, molluscs, fish, frogs, and small rodents[95]
 LC 


Unknown Unknown[95]

Genus Notiosorex Coues, 1877 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Cockrum's gray shrew

Gray shrew specimen

N. cockrumi
Baker, O'Neill, & McAliley, 2003
Northern Mexico and southern United States
Map of range
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Shrubland and desert[97]

Diet: Insects and other invertebrates, as well as carrion[87]
 LC 


Unknown Population steady[97]

Crawford's gray shrew

Gray shrew

N. crawfordi
(Coues, 1877)
Southern North America
Map of range
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Forest, shrubland, grassland, and desert[98]

Diet: Worms, spiders, and insects, as well as small mammals, lizards, and birds[99]
 LC 


Unknown Population steady[98]

Large-eared gray shrew


N. evotis
(Coues, 1877)
Western Mexico
Map of range
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Shrubland and grassland[100]

Diet: Insects and other invertebrates, as well as carrion[87]
 LC 


Unknown Unknown[100]

Villa's gray shrew


N. villai
Carraway & Timm, 2000
Eastern Mexico
Map of range
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Forest[101]

Diet: Insects and other invertebrates, as well as carrion[87]
 VU 


Unknown Population declining[101]

Genus Sorex Linnaeus, 1758 – 76 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Alaska tiny shrew


S. yukonicus
Dokuchaev, 1997
Alaska
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands[103]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[103]

Alpine shrew


S. alpinus
Schinz, 1837
Europe
Map of range
Size: 6–8 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail[105]

Habitat: Forest, grassland, and rocky areas[106]

Diet: Insects, arachnids, snails, and earthworms[105]
 NT 


Unknown Population declining[106]

American pygmy shrew


S. hoyi
Baird, 1857
Northern North America
Map of range
Size: 5–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail[107]

Habitat: Forest, shrubland, grassland, and inland wetlands[108]

Diet: Insects and other invertebrates[107]
 LC 


Unknown Population steady[108]

American water shrew

Brown shrew

S. palustris
Richardson, 1828
North America
Map of range
Size: 7–9 cm (3–4 in) long, plus 5–9 cm (2–4 in) tail[109]

Habitat: Forest and inland wetlands[110]

Diet: Aquatic insects and fish, as well as other invertebrates and plants[109]
 LC 


Unknown Population steady[110]

Apennine shrew


S. samniticus
Altobello, 1926
Italy
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland and forest[111]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[111]

Arctic shrew


S. arcticus
Kerr, 1792
Northern North America
Map of range
Size: 6–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[112]

Habitat: Forest and inland wetlands[113]

Diet: Insects and other invertebrates[112]
 LC 


Unknown Population steady[113]

Arizona shrew


S. arizonae
Diersing & Hoffmeister, 1977
Northern Mexico and southern United States
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[114]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[114]

Azumi shrew


S. hosonoi
Imaizumi, 1954
Japan
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and grassland[115]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[115]

Baird's shrew

Brown shrew

S. bairdi
Merriam, 1895
Northwestern United States
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[116]

Diet: Invertebrates[116]
 LC 


Unknown Unknown[116]

Barren ground shrew


S. ugyunak
Anderson & Rand, 1945
Northern North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland[117]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[117]

Buchara shrew

Stamp with brown shrew

S. buchariensis
Ognew, 1921
Central Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[118]

Diet: Insects[118]
 DD 


Unknown Unknown[118]

Carmen Mountain shrew


S. milleri
Jackson, 1947
Northeastern Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[119]

Diet: Insects[119]
 VU 


Unknown Population declining[119]

Caucasian pygmy shrew

Brown shrew

S. volnuchini
Ognew, 1922
Western Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[120]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[120]

Caucasian shrew


S. satunini
Ognew, 1922
Western Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and rocky areas[121]

Diet: Insects[121]
 LC 


Unknown Unknown[121]

Chestnut-bellied shrew


S. ventralis
Merriam, 1895
Southern Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[122]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[122]

Chinese highland shrew


S. excelsus
Allen, 1923
Southern China
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and grassland[123]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[123]

Chinese shrew


S. sinalis
Thomas, 1912
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Rocky areas[124]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[124]

Cinereus shrew

Brown shrew

S. cinereus
Kerr, 1792
Northern North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, and inland wetlands[125]

Diet: A variety of invertebrates, as well as seeds and fungi[126]
 LC 


Unknown Population steady[125]

Common shrew

Brown shrew

S. araneus
Linnaeus, 1758
Europe and Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, inland wetlands, and coastal marine[127]

Diet: Insects and other invertebrates[128]
 LC 


Unknown Population steady[127]

Crowned shrew

Brown shrew

S. coronatus
Millet, 1828
Western Europe
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and inland wetlands[129]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[129]

Dwarf shrew


S. nanus
Merriam, 1895
Central United States
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Rocky areas, inland wetlands, grassland, and forest[130]

Diet: Insects and other invertebrates, as well as plants and small vertebrates[130]
 LC 


Unknown Unknown[130]

Eurasian least shrew

Brown shrew

S. minutissimus
Zimmermann, 1780
Norther Europe and northern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[131]

Diet: Insects and other invertebrates[131]
 LC 


Unknown Unknown[131]

Eurasian pygmy shrew

Brown shrew

S. minutus
Linnaeus, 1766
Europe and Asia
Map of range
Size: 3–6 cm (1–2 in) long, plus 3–5 cm (1–2 in) tail[132]

Habitat: Forest, shrubland, grassland, inland wetlands, and coastal marine[133]

Diet: Insects and other invertebrates[132]
 LC 


Unknown Population steady[133]

Flat-skulled shrew


S. roboratus
Hollister, 1913
Northern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[134]

Diet: Insects[134]
 LC 


Unknown Unknown[134]

Fog shrew


S. sonomae
Jackson, 1921

Two subspecies
  • S. s. sonomae
  • S. s. tenelliodus
Western North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[135]

Diet: Insects and other invertebrates[135]
 LC 


Unknown Population steady[135]

Gansu shrew


S. cansulus
Thomas, 1912
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Unknown[136]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[136]

Glacier Bay water shrew


S. alaskanus
Merriam, 1900
Western Canada
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands[137]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[137]

Greater stripe-backed shrew


S. cylindricauda
H. Milne-Edwards, 1872
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[138]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[138]

Iberian shrew

Brown shrew

S. granarius
Miller, 1910
Portugal and Spain
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and rocky areas[139]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[139]

Inyo shrew

Drawing of shrew skulls

S. tenellus
Merriam, 1895
Western United States
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, inland wetlands, and rocky areas[140]

Diet: Insects and other invertebrates[140]
 LC 


Unknown Population steady[140]

Ixtlan shrew


S. ixtlanensis
Carraway, 2007
Southern Mexico Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[141]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[141]

Jalisco shrew


S. mediopua
Carraway, 2007
Southern Mexico Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[142]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[142]

Kamchatka shrew


S. camtschatica
Yudin, 1972
Eastern Russia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland[143]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[143]

Kashmir pygmy shrew


S. planiceps
Miller, 1911
Northern India and Pakistan
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and rocky areas[144]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[144]

Kozlov's shrew


S. kozlovi
Stroganov, 1952
South-central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Unknown[145]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[145]

Large-toothed shrew


S. macrodon
Merriam, 1895
Southern Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[146]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 VU 


Unknown Population declining[146]

Laxmann's shrew

Brown shrew

S. caecutiens
Laxmann, 1788
Northern Europe and northern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and inland wetlands[147]

Diet: Insects and other invertebrates[147]
 LC 


Unknown Population steady[147]

Lesser striped shrew


S. bedfordiae
Thomas, 1911
Central China and southern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[148]

Diet: Insects[148]
 LC 


Unknown Unknown[148]

Long-clawed shrew

Brown shrew

S. unguiculatus
Dobson, 1890
Eastern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[149]

Diet: Insects and other invertebrates[149]
 LC 


Unknown Population steady[149]

Long-tailed shrew

Brown shrew

S. dispar
Batchelder, 1911
Eastern North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[150]

Habitat: Forest, inland wetlands, and rocky areas[151]

Diet: Small invertebrates and plants[150]
 LC 


Unknown Unknown[151]

Maritime shrew


S. maritimensis
Smith, 1939
Eastern Canada
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, inland wetlands, and unknown[152]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population declining[152]

Marsh shrew

Black shrew

S. bendirii
Merriam, 1884
Western North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[153]

Diet: Insects[153]
 LC 


Unknown Unknown[153]

Merriam's shrew

Gray shrew

S. merriami
Dobson, 1890
Western United States
Map of range
Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[154]

Habitat: Forest, shrubland, and grassland[155]

Diet: Insects and other invertebrates, as well as small vertebrates[154]
 LC 


Unknown Unknown[155]

Mexican long-tailed shrew


S. oreopolus
Merriam, 1892
Southern Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[156]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[156]

Montane shrew

Brown shrew

S. monticolus
Merriam, 1890
Western North America
Map of range
Size: 6–8 cm (2–3 in) long, plus 4–7 cm (2–3 in) tail[157]

Habitat: Forest, grassland, and inland wetlands[158]

Diet: Insects and other invertebrates, as well as salamanders, seeds, lichen, and fungi[157]
 LC 


Unknown Population steady[158]

Mount Lyell shrew


S. lyelli
Merriam, 1902
Western United States
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland, grassland, and inland wetlands[159]

Diet: Insects[159]
 LC 


Unknown Population steady[159]

New Mexico shrew


S. neomexicanus
Bailey, 1913
Southern United States
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[160]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[160]

Orizaba long-tailed shrew


S. orizabae
Merriam, 1895
Central Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[161]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[161]

Ornate shrew

Gray shrew

S. ornatus
Merriam, 1895

Eight subspecies
  • S. o. juncensis (Tule shrew)
  • S. o. lagunae
  • S. o. ornatus
  • S. o. relictus
  • S. o. salarius
  • S. o. salicornicus
  • S. o. willetti
  • S. o. sinuosus (Suisun shrew)
Western North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, and inland wetlands[162]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[162]

Pacific shrew


S. pacificus
Coues, 1877

Two subspecies
  • S. p. cascadensis
  • S. p. pacificus
Western United States
Map of range
Size: 8–9 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail[163]

Habitat: Forest[164]

Diet: Insects, other invertebrates, and amphibians, as well as plants and fungi[163]
 LC 


Unknown Population steady[164]

Paramushir shrew


S. leucogaster
Kuroda, 1933
Eastern Russia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland[165]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[165]

Portenko's shrew


S. portenkoi
Stroganov, 1956
Eastern Russia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland[166]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[166]

Prairie shrew


S. haydeni
Baird, 1857
Central North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, and inland wetlands[167]

Diet: Insects and other invertebrates, as well as small vertebrates and plants[167]
 LC 


Unknown Population steady[167]

Preble's shrew


S. preblei
Jackson, 1922
Northwestern United States
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, inland wetlands, and desert[168]

Diet: Insects[168]
 LC 


Unknown Unknown[168]

Pribilof Island shrew


S. pribilofensis
Merriam, 1895
Pribilof Islands in Alaska
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland and coastal marine[169]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 EN 


Unknown Unknown[169]

Radde's shrew


S. raddei
Satunin, 1895
Western Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and rocky areas[170]

Diet: Insects[170]
 LC 


Unknown Unknown[170]

Saint Lawrence Island shrew


S. jacksoni
Hall & Gilmore, 1932
St. Lawrence Island in Alaska
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland, inland wetlands, and rocky areas[171]

Diet: Insects[171]
 LC 


Unknown Unknown[171]

San Cristobal shrew


S. stizodon
Merriam, 1895
Southeastern Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[172]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 CR 


Unknown Population declining[172]

Saussure's shrew


S. saussurei
Merriam, 1892
Southern Mexico and Guatemala
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[173]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[173]

Sclater's shrew


S. sclateri
Merriam, 1897
Southern Mexico and Guatemala
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[174]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 CR 


Unknown Population declining[174]

Shinto shrew


S. shinto
Thomas, 1905

Three subspecies
  • S. s. sadonis (Sado shrew)
  • S. s. shikokensis
  • S. s. shinto
Japan
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[175]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[175]

Siberian large-toothed shrew


S. daphaenodon
Thomas, 1907
Northern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands and forest[176]

Diet: Insects and other invertebrates[176]
 LC 


Unknown Population steady[176]

Slender shrew


S. gracillimus
Thomas, 1907

Four subspecies
  • S. g. gracillimus
  • S. g. granti
  • S. g. minor
  • S. g. natalae
Eastern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[177]

Diet: Invertebrates[177]
 LC 


Unknown Population steady[177]

Smoky shrew

Gray shrew

S. fumeus
Miller, 1895
Eastern North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[178]

Diet: Invertebrates, as well as salamanders and fungi[179]
 LC 


Unknown Population steady[178]

Southeastern shrew

Gray shrew

S. longirostris
Bachman, 1837

Three subspecies
  • S. l. eonis
  • S. l. fisheri (Dismal Swamp southeastern shrew)
  • S. l. longirostris
Eastern United States
Map of range
Size: 4–6 cm (2–2 in) long, plus 2–4 cm (1–2 in) tail[180]

Habitat: Inland wetlands, grassland, shrubland, and forest[181]

Diet: Insects and other invertebrates as well as seeds[180]
 LC 


Unknown Population steady[181]

Taiga shrew


S. isodon
Turov, 1924
Northern Europe and northern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[182]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[182]

Tibetan shrew


S. thibetanus
Kaschtschenko, 1905
Central China
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and grassland[183]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[183]

Tien Shan shrew


S. asper
Thomas, 1914
Central Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[184]

Diet: Insects and other invertebrates[184]
 LC 


Unknown Unknown[184]

Trowbridge's shrew

Gray shrew

S. trowbridgii
Baird, 1857
Western North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and inland wetlands[185]

Diet: Insects and other invertebrates[185]
 LC 


Unknown Population steady[185]

Tundra shrew

Gray shrew

S. tundrensis
Merriam, 1900
Asia and western North America
Map of range
Size: 8–12 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[186]

Habitat: Forest, grassland, and inland wetlands[187]

Diet: Insects, other invertebrates, and flowers[186]
 LC 


Unknown Population steady[187]

Ussuri shrew


S. mirabilis
Ognew, 1937
Eastern Asia
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[188]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown Unknown[188]

Vagrant shrew

Brown shrew

S. vagrans
Baird, 1857
Western North America
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[189]

Diet: Insects and other invertebrates[189]
 LC 


Unknown Population steady[189]

Valais shrew

Brown shrew

S. antinorii
Bonaparte, 1840
Southern Europe
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands[190]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Unknown[190]

Veracruz shrew


S. veraecrucis
Jackson, 1925
Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[191]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[191]

Verapaz shrew


S. veraepacis
Alston, 1877

Three subspecies
  • S. v. chiapensis
  • S. v. mutabilis
  • S. v. veraepacis
Southern Mexico and Guatemala
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[192]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population steady[192]

Zacatecas shrew


S. emarginatus
Jackson, 1925
Central Mexico
Map of range
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[193]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown Population declining[193]

Genus Soriculus Blyth, 1854 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Himalayan shrew

Brown shrew

S. nigrescens
(Gray, 1842)
Himalayas in Asia
Map of range
Size: 8–10 cm (3–4 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest, shrubland, and rocky areas[194]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown Unknown[194]

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