Phoronids (scientific name Phoronida, sometimes called horseshoe worms) are a small phylum of marine animals that filter-feed with a lophophore (a "crown" of tentacles), and build upright tubes of chitin to support and protect their soft bodies. They live in most of the oceans and seas, including the Arctic Ocean but excluding the Antarctic Ocean, and between the intertidal zone and about 400 meters down. Most adult phoronids are 2 cm long and about 1.5 mm wide, although the largest are 50 cm long.
Livyatan is an extinctgenus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. Herman Melville often referred to whales as "Leviathans" in his book. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Mioceneepoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, the United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales (or "raptorial sperm whales") and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.
Livyatan's total length has been estimated to be about 13.5–17.5 m (44–57 ft), almost similar to that of the modern sperm whale (Physeter macrocephalus), making it one of the largest predators known to have existed. The teeth of Livyatan measured 36.2 cm (1.19 ft), and are the largest biting teeth of any known animal, excluding tusks. It is distinguished from the other raptorial sperm whales by the basin on the skull spanning the length of the snout. The spermaceti organ contained in that basin is thought to have been used in echolocation and communication, or for ramming prey and other sperm whales. The whale may have interacted with the large extinct shark megalodon (Otodus megalodon), competing with it for a similar food source. Its extinction was probably caused by a cooling event at the end of the Miocene period causing a reduction in food populations. The geological formation where the whale has been found has also preserved a large assemblage of marine life, such as sharks and marine mammals. (Full article...)
C. maenas is a widespread invasive species, listed among the 100 of the World's Worst Invasive Alien Species. It is native to the north-east Atlantic Ocean and Baltic Sea, but has colonised similar habitats in Australia, South Africa, South America and both Atlantic and Pacific coasts of North America. It grows to a carapace width of 90 mm (3+1⁄2 in), and feeds on a variety of mollusks, worms, and small crustaceans, affecting a number of fisheries. Its successful dispersal has occurred by a variety of mechanisms, such as on ships' hulls, sea planes, packing materials, and bivalves moved for aquaculture. (Full article...)
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Pinnipedia is an infraorder of mammals in the orderCarnivora, composed of seals, sea lions, and the walrus. A member of this group is called a pinniped or a seal. They are widespread throughout the ocean and some larger lakes, primarily in colder waters. Pinnipeds range in size from the 1.1 m (3 ft 7 in) and 50 kg (110 lb) Baikal seal to the 6 m (20 ft) and 3,700 kg (8,200 lb) male southern elephant seal, which is also the largest member of Carnivora. Several species exhibit sexual dimorphism, such as the southern elephant seal, where the males can be more than three times as long and six times as massive as the females, or the Ross seal, which has females typically larger than the males. Four seal species are estimated to have over one million members, while six are classified as endangered with population counts as low as 600, and two, the Caribbean monk seal and the Japanese sea lion, went extinct in the 20th century.
The 34 extant species of Pinnipedia are split into 22 genera within 3 families: Odobenidae, comprising the walrus; Otariidae, the eared seals, split between the sea lions and fur seals; and Phocidae, the earless or true seals. Odobenidae and Otariidae are combined into the superfamilyOtarioidea, with Phocidae in Phocoidea. Extinct species have also been placed into the three extant families, as well as the extinct family Desmatophocidae, though most extinct species have not been categorized into a subfamily. Nearly one hundred extinct Pinnipedia species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed. (Full article...)
Marine mammal adaptation to an aquatic lifestyle varies considerably between species. Both cetaceans and sirenians are fully aquatic and therefore are obligate water dwellers. Pinnipeds are semiaquatic; they spend the majority of their time in the water but need to return to land for important activities such as mating, breeding and molting. Sea otters tend to live in kelp forests and estuaries. In contrast, the polar bear is mostly terrestrial and only go into the water on occasions of necessity, and are thus much less adapted to aquatic living. The diets of marine mammals vary considerably as well; some eat zooplankton, others eat fish, squid, shellfish, or seagrass, and a few eat other mammals. While the number of marine mammals is small compared to those found on land, their roles in various ecosystems are large, especially concerning the maintenance of marine ecosystems, through processes including the regulation of prey populations. This role in maintaining ecosystems makes them of particular concern as 23% of marine mammal species are currently threatened. (Full article...)
Manta rays are large rays belonging to the genusMobula (formerly its own genusManta). The larger species, M. birostris, reaches 7 m (23 ft) in width, while the smaller, M. alfredi, reaches 5.5 m (18 ft). Both have triangular pectoral fins, horn-shaped cephalic fins and large, forward-facing mouths. They are classified among the Myliobatiformes (stingrays and relatives) and are placed in the familyMyliobatidae (eagle rays). They have the largest brain-to-body ratio of all fish, and can pass the mirror test.
Mantas are found in warm temperate, subtropical and tropical waters. Both species are pelagic; M. birostrismigrates across open oceans, singly or in groups, while M. alfredi tends to be resident and coastal. They are filter feeders and eat large quantities of zooplankton, which they gather with their open mouths as they swim. However, research suggests that the majority of their diet (73%) comes from mesopelagic sources. Gestation lasts over a year and mantas give birth to live pups. Mantas may visit cleaning stations for the removal of parasites. Like whales, they breach for unknown reasons. (Full article...)
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Steller's sea ape is a purported marine mammal, observed by German zoologist Georg Steller on August 10, 1741, around the Shumagin Islands in Alaska. The animal was described as being around 1.5 m (5 feet) long; with a dog-like head; long drooping whiskers; an elongated but robust body; thick fur coat; no limbs; and tail fins much like a shark. He described the creature as being playful and inquisitive like a monkey. After observing it for two hours, he attempted to shoot and collect the creature, but missed, and the creature swam away.
There have been four attempts to scientifically classify the creature, described as Simia marina, Siren cynocephala, Trichechus hydropithecus, and Manatus simia. Most likely, Steller simply misidentified a northern fur seal. (Full article...)
Terropterus was the earliest known and largest mixopterid eurypterid. Fossil specimens referred to T. xiushanensis are estimated to have reached up to 40 centimeters (15.7 in) in length, but other fossils, either representing older T. xiushanensis or a second species of Terropterus, demonstrate that members of the genus could reach upwards of at least 100 centimeters (3.3 ft) in length. Terropterus is the only mixopterid known from the ancient southern continent of Gondwana, with the other two mixopterid genera, Mixopterus and Lanarkopterus, only being known from what was once the northern continent of Laurussia. The discovery of Terropterus significantly expanded the known geographical and temporal ranges of the Mixopteridae. (Full article...)
Teleostei (/ˌtɛliˈɒstiaɪ/; Greekteleios "complete" + osteon "bone"), members of which are known as teleosts (/ˈtɛliɒsts,ˈtiːli-/), is, by far, the largest infraclass in the class Actinopterygii, the ray-finned fishes, and contains 96% of all extant species of fish. Teleosts are arranged into about 40 orders and 448 families. Over 26,000 species have been described. Teleosts range from giant oarfish measuring 7.6 m (25 ft) or more, and ocean sunfish weighing over 2 t (2.0 long tons; 2.2 short tons), to the minute male anglerfishPhotocorynus spiniceps, just 6.2 mm (0.24 in) long. Including not only torpedo-shaped fish built for speed, teleosts can be flattened vertically or horizontally, be elongated cylinders or take specialised shapes as in anglerfish and seahorses.
The difference between teleosts and other bony fish lies mainly in their jaw bones; teleosts have a movable premaxilla and corresponding modifications in the jaw musculature which make it possible for them to protrude their jaws outwards from the mouth. This is of great advantage, enabling them to grab prey and draw it into the mouth. In more derived teleosts, the enlarged premaxilla is the main tooth-bearing bone, and the maxilla, which is attached to the lower jaw, acts as a lever, pushing and pulling the premaxilla as the mouth is opened and closed. Other bones further back in the mouth serve to grind and swallow food. Another difference is that the upper and lower lobes of the tail (caudal) fin are about equal in size. The spine ends at the caudal peduncle, distinguishing this group from other fish in which the spine extends into the upper lobe of the tail fin. (Full article...)
Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the cladeSelachimorpha (or Selachii) and are the sister group to the Batoidea (rays and kin). Some sources extend the term "shark" as an informal category including extinct members of Chondrichthyes (cartilaginous fish) with a shark-like morphology, such as hybodonts. Shark-like chondrichthyans such as Cladoselache and Doliodus first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as the Late Ordovician (458–444 million years ago). The earliest confirmed modern sharks (selachimorphs) are known from the Early Jurassic around 200 million years ago, with the oldest known member being Agaleus, though records of true sharks may extend back as far as the Permian.
Sharks range in size from the small dwarf lanternshark (Etmopterus perryi), a deep sea species that is only 17 centimetres (6.7 in) in length, to the whale shark (Rhincodon typus), the largest fish in the world, which reaches approximately 12 metres (40 ft) in length. They are found in all seas and are common to depths up to 2,000 metres (6,600 ft). They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river sharks, which can be found in both seawater and freshwater, and the Ganges shark, which lives only in freshwater. Sharks have a covering of dermal denticles that protects their skin from damage and parasites in addition to improving their fluid dynamics. They have numerous sets of replaceable teeth. (Full article...)
Image 2A microbial mat encrusted with iron oxide on the flank of a seamount can harbour microbial communities dominated by the iron-oxidizing Zetaproteobacteria (from Marine prokaryotes)
Image 4Sandy shores provide shifting homes to many species (from Marine habitat)
Image 5Marine Species Changes in Latitude and Depth in three different ocean regions(1973–2019) (from Marine food web)
Image 6Antarctic marine food web. Potter Cove 2018. Vertical position indicates trophic level and node widths are proportional to total degree (in and out). Node colors represent functional groups. (from Marine food web)
Image 7Estuaries occur when rivers flow into a coastal bay or inlet. They are nutrient rich and have a transition zone which moves from freshwater to saltwater. (from Marine habitat)
Image 10The Ocean Cleanup is one of many organizations working toward marine conservation such at this interceptor vessel that prevents plastic from entering the ocean. (from Marine conservation)
Image 11Elevation-area graph showing the proportion of land area at given heights and the proportion of ocean area at given depths (from Marine habitat)
Image 12
Diagram of a mycoloop (fungus loop)
Parasitic chytrids can transfer material from large inedible phytoplankton to zooplankton. Chytrids zoospores are excellent food for zooplankton in terms of size (2–5 μm in diameter), shape, nutritional quality (rich in polyunsaturated fatty acids and cholesterols). Large colonies of host phytoplankton may also be fragmented by chytrid infections and become edible to zooplankton. (from Marine fungi)
Image 16Oceanic pelagic food web showing energy flow from micronekton to top predators. Line thickness is scaled to the proportion in the diet. (from Marine food web)
Image 18Conference events, such as the events hosted by the United Nations, help to bring together many stakeholders for awareness and action. (from Marine conservation)
Image 19Scanning electron micrograph of a strain of Roseobacter, a widespread and important genus of marine bacteria. For scale, the membrane pore size is 0.2μm in diameter. (from Marine prokaryotes)
Image 20Tidepools on rocky shores make turbulent habitats for many forms of marine life (from Marine habitat)
Image 21Anthropogenic stressors to marine species threatened with extinction (from Marine food web)
Image 23Cryptic interactions in the marine food web. Red: mixotrophy; green: ontogenetic and species differences; purple: microbial cross‐feeding; orange: auxotrophy; blue: cellular carbon partitioning. (from Marine food web)
Image 24Schematic representation of the changes in abundance between trophic groups in a temperate rocky reef ecosystem. (a) Interactions at equilibrium. (b) Trophic cascade following disturbance. In this case, the otter is the dominant predator and the macroalgae are kelp. Arrows with positive (green, +) signs indicate positive effects on abundance while those with negative (red, -) indicate negative effects on abundance. The size of the bubbles represents the change in population abundance and associated altered interaction strength following disturbance. (from Marine food web)
Image 25Phylogenetic and symbiogenetic tree of living organisms, showing a view of the origins of eukaryotes and prokaryotes (from Marine prokaryotes)
Image 33Coral reefs provide marine habitats for tube sponges, which in turn become marine habitats for fishes (from Marine habitat)
Image 34Reconstruction of an ammonite, a highly successful early cephalopod that first appeared in the Devonian (about 400 mya). They became extinct during the same extinction event that killed the land dinosaurs (about 66 mya). (from Marine invertebrates)
Image 41Ernst Haeckel's 96th plate, showing some marine invertebrates. Marine invertebrates have a large variety of body plans, which are currently categorised into over 30 phyla. (from Marine invertebrates)
Image 43Cnidarians are the simplest animals with cells organised into tissues. Yet the starlet sea anemone contains the same genes as those that form the vertebrate head. (from Marine invertebrates)
Image 44An in situ perspective of a deep pelagic food web derived from ROV-based observations of feeding, as represented by 20 broad taxonomic groupings. The linkages between predator to prey are coloured according to predator group origin, and loops indicate within-group feeding. The thickness of the lines or edges connecting food web components is scaled to the log of the number of unique ROV feeding observations across the years 1991–2016 between the two groups of animals. The different groups have eight colour-coded types according to main animal types as indicated by the legend and defined here: red, cephalopods; orange, crustaceans; light green, fish; dark green, medusa; purple, siphonophores; blue, ctenophores and grey, all other animals. In this plot, the vertical axis does not correspond to trophic level, because this metric is not readily estimated for all members. (from Marine food web)
Different bacteria shapes (cocci, rods and spirochetes) and their sizes compared with the width of a human hair. A few bacteria are comma-shaped (vibrio). Archaea have similar shapes, though the archaeon Haloquadratum is flat and square.
The unit μm is a measurement of length, the micrometer, equal to 1/1,000 of a millimeter
Image 52A protected sea turtle area that warns of fines and imprisonment on a beach in Miami, Florida. (from Marine conservation)
Image 53Halfbeak as larvae are one of the organisms adapted to the unique properties of the microlayer (from Marine habitat)
Image 54Conceptual diagram of faunal community structure and food-web patterns along fluid-flux gradients within Guaymas seep and vent ecosystems. (from Marine food web)
Image 55Ocean surface chlorophyll concentrations in October 2019. The concentration of chlorophyll can be used as a proxy to indicate how many phytoplankton are present. Thus on this global map green indicates where a lot of phytoplankton are present, while blue indicates where few phytoplankton are present. – NASA Earth Observatory 2019. (from Marine food web)
Image 56The distribution of anthropogenic stressors faced by marine species threatened with extinction in various marine regions of the world. Numbers in the pie charts indicate the percentage contribution of an anthropogenic stressors' impact in a specific marine region. (from Marine food web)
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Mycoloop links between phytoplankton and zooplankton
Chytrid‐mediated trophic links between phytoplankton and zooplankton (mycoloop). While small phytoplankton species can be grazed upon by zooplankton, large phytoplankton species constitute poorly edible or even inedible prey. Chytrid infections on large phytoplankton can induce changes in palatability, as a result of host aggregation (reduced edibility) or mechanistic fragmentation of cells or filaments (increased palatability). First, chytrid parasites extract and repack nutrients and energy from their hosts in form of readily edible zoospores. Second, infected and fragmented hosts including attached sporangia can also be ingested by grazers (i.e. concomitant predation). (from Marine fungi)
Image 59Phylogenetic tree representing bacterial OTUs from clone libraries and next-generation sequencing. OTUs from next-generation sequencing are displayed if the OTU contained more than two sequences in the unrarefied OTU table (3626 OTUs). (from Marine prokaryotes)
Image 60Jellyfish are easy to capture and digest and may be more important as food sources than was previously thought. (from Marine food web)
Image 61Whales were close to extinction until legislation was put in place. (from Marine conservation)
Image 66Dickinsonia may be the earliest animal. They appear in the fossil record 571 million to 541 million years ago. (from Marine invertebrates)
Image 67Sea ice food web and the microbial loop. AAnP = aerobic anaerobic phototroph, DOC = dissolved organic carbon, DOM = dissolved organic matter, POC = particulate organic carbon, PR = proteorhodopsins. (from Marine food web)
Image 68Biomass pyramids. Compared to terrestrial biomass pyramids, aquatic pyramids are generally inverted at the base. (from Marine food web)
Image 69Common-enemy graph of Antarctic food web. Potter Cove 2018. Nodes represent basal species and links indirect interactions (shared predators). Node and link widths are proportional to number of shared predators. Node colors represent functional groups. (from Marine food web)
Image 70The deep sea amphipodEurythenes plasticus, named after microplastics found in its body, demonstrating plastic pollution affects marine habitats even 6000m below sea level. (from Marine habitat)
Image 71640 μm microplastic found in the deep sea amphipod Eurythenes plasticus (from Marine habitat)
Image 75Some representative ocean animal life (not drawn to scale) within their approximate depth-defined ecological habitats. Marine microorganisms exist on the surfaces and within the tissues and organs of the diverse life inhabiting the ocean, across all ocean habitats. (from Marine habitat)
Image 84Only 29 percent of the world surface is land. The rest is ocean, home to the marine habitats. The oceans are nearly four kilometres deep on average and are fringed with coastlines that run for nearly 380,000 kilometres.
Image 86On average there are more than one million microbial cells in every drop of seawater, and their collective metabolisms not only recycle nutrients that can then be used by larger organisms but also catalyze key chemical transformations that maintain Earth's habitability. (from Marine food web)
Image 88Food web structure in the euphotic zone. The linear food chain large phytoplankton-herbivore-predator (on the left with red arrow connections) has fewer levels than one with small phytoplankton at the base. The microbial loop refers to the flow from the dissolved organic carbon (DOC) via heterotrophic bacteria (Het. Bac.) and microzooplankton to predatory zooplankton (on the right with black solid arrows). Viruses play a major role in the mortality of phytoplankton and heterotrophic bacteria, and recycle organic carbon back to the DOC pool. Other sources of dissolved organic carbon (also dashed black arrows) includes exudation, sloppy feeding, etc. Particulate detritus pools and fluxes are not shown for simplicity. (from Marine food web)
Image 89In the open ocean, sunlit surface epipelagic waters get enough light for photosynthesis, but there are often not enough nutrients. As a result, large areas contain little life apart from migrating animals. (from Marine habitat)
Image 91A 2016 metagenomic representation of the tree of life using ribosomal protein sequences. The tree includes 92 named bacterial phyla, 26 archaeal phyla and five eukaryotic supergroups. Major lineages are assigned arbitrary colours and named in italics with well-characterized lineage names. Lineages lacking an isolated representative are highlighted with non-italicized names and red dots. (from Marine prokaryotes)
Image 95This algae bloom occupies sunlit epipelagic waters off the southern coast of England. The algae are maybe feeding on nutrients from land runoff or upwellings at the edge of the continental shelf. (from Marine habitat)
Estimates of microbial species counts in the three domains of life
Bacteria are the oldest and most biodiverse group, followed by Archaea and Fungi (the most recent groups). In 1998, before awareness of the extent of microbial life had gotten underway, Robert M. May estimated there were 3 million species of living organisms on the planet. But in 2016, Locey and Lennon estimated the number of microorganism species could be as high as 1 trillion. (from Marine prokaryotes)
Image 99Some lobe-finned fishes, like the extinct Tiktaalik, developed limb-like fins that could take them onto land (from Marine vertebrate)
Image 102Topological positions versus mobility: (A) bottom-up groups (sessile and drifters), (B) groups at the top of the food web. Phyto, phytoplankton; MacroAlga, macroalgae; Proto, pelagic protozoa; Crus, Crustacea; PelBact, pelagic bacteria; Echino, Echinoderms; Amph, Amphipods; HerbFish, herbivorous fish; Zoopl, zooplankton; SuspFeed, suspension feeders; Polych, polychaetes; Mugil, Mugilidae; Gastropod, gastropods; Blenny, omnivorous blennies; Decapod, decapods; Dpunt, Diplodus puntazzo; Macropl, macroplankton; PlFish, planktivorous fish; Cephalopod, cephalopods; Mcarni, macrocarnivorous fish; Pisc, piscivorous fish; Bird, seabirds; InvFeed1 through InvFeed4, benthic invertebrate feeders. (from Marine food web)
Image 103The pelagic food web, showing the central involvement of marine microorganisms in how the ocean imports nutrients from and then exports them back to the atmosphere and ocean floor (from Marine food web)
Solar radiation can have positive (+) or negative (−) effects resulting in increases or decreases in the heterotrophic activity of bacterioplankton. (from Marine prokaryotes)
Model of the energy generating mechanism in marine bacteria
(1) When sunlight strikes a rhodopsin molecule (2) it changes its configuration so a proton is expelled from the cell (3) the chemical potential causes the proton to flow back to the cell (4) thus generating energy (5) in the form of adenosine triphosphate. (from Marine prokaryotes)
Image 110Chytrid parasites of marine diatoms. (A) Chytrid sporangia on Pleurosigma sp. The white arrow indicates the operculate discharge pore. (B) Rhizoids (white arrow) extending into diatom host. (C) Chlorophyll aggregates localized to infection sites (white arrows). (D and E) Single hosts bearing multiple zoosporangia at different stages of development. The white arrow in panel E highlights branching rhizoids. (F) Endobiotic chytrid-like sporangia within diatom frustule. Bars = 10 μm. (from Marine fungi)
Image 111Cycling of marine phytoplankton. Phytoplankton live in the photic zone of the ocean, where photosynthesis is possible. During photosynthesis, they assimilate carbon dioxide and release oxygen. If solar radiation is too high, phytoplankton may fall victim to photodegradation. For growth, phytoplankton cells depend on nutrients, which enter the ocean by rivers, continental weathering, and glacial ice meltwater on the poles. Phytoplankton release dissolved organic carbon (DOC) into the ocean. Since phytoplankton are the basis of marine food webs, they serve as prey for zooplankton, fish larvae and other heterotrophic organisms. They can also be degraded by bacteria or by viral lysis. Although some phytoplankton cells, such as dinoflagellates, are able to migrate vertically, they are still incapable of actively moving against currents, so they slowly sink and ultimately fertilize the seafloor with dead cells and detritus. (from Marine food web)
Image 112Waves and currents shape the intertidal shoreline, eroding the softer rocks and transporting and grading loose particles into shingles, sand or mud (from Marine habitat)
Image 113This timeline contains clickable links
Image 114Ocean or marine biomass, in a reversal of terrestrial biomass, can increase at higher trophic levels. (from Marine food web)
Image 121Archaea were initially viewed as extremophiles living in harsh environments, such as the yellow archaea pictured here in a hot spring, but they have since been found in a much broader range of habitats. (from Marine prokaryotes)
Image 4Global map of large marine ecosystems. Oceanographers and biologists have identified 66 LMEs worldwide. (from Marine ecosystem)
Image 5Ecosystem services delivered by epibenthicbivalve reefs. Reefs provide coastal protection through erosion control and shoreline stabilization, and modify the physical landscape by ecosystem engineering, thereby providing habitat for species by facilitative interactions with other habitats such as tidal flat benthic communities, seagrasses and marshes. (from Marine ecosystem)
... because whales and dolphins are streamlined to swim in water, they do not have external organs. This makes it almost impossible to tell the sex of a whale or dolphin when watching them on the sea surface.
... If sharks don’t keep on swimming they sink to the seabed.
... Shark brains aren’t round like a human's; they are long and narrow.
... in baleen whales females are bigger than males, while in the toothed whales the males are bigger than females.
... A shark can sense a turtle, octopus or other prey from up to 20m away.
... Most sharks never close their eyes. Some have special see-through eyelids that protect their eyes without cutting out light. Others just roll their eyes up into their head to protect them.
Cuttlefish are sometimes called the chameleon of the sea because of their remarkable ability to rapidly alter their skin colour at will. Their skin flashes a fast-changing pattern as communication to other cuttlefish and to camouflage them from predators.