Hunsrück Slate
| Hunsrück Slate | |
|---|---|
| Stratigraphic range: latest Pragian to early Emsian | |
 X-ray photograph of fossil of Chotecops ferdianandi, a common trilobite from Hunsrück Slate | |
| Type | Geological formation |
| Lithology | |
| Primary | Slate |
| Location | |
| Region | Hunsrück |
| Country | Germany |
The Hunsrück Slate (German: Hunsrück-Schiefer) is a Lower Devonian lithostratigraphic unit, a type of rock strata, in the German regions of the Hunsrück and Taunus. It is a lagerstätte famous for exceptional preservation of a highly diverse fossil fauna assemblage.
Geology
The Emsian stratigraphy of the southern Rhenish Massif can be divided into two lithological units: the older slates of the Hunsrück-Schiefer and the younger sandstones of the Singhofener Schichten. Stratigraphically below the Hunsrück Slates is the (older) Taunus quartzite.[1] All these metasedimentary rocks were originally deposited in the marine Rhenohercynian Basin, a back-arc basin south of the paleocontinent of Laurussia.
The Hunsrück Slate roughly comprises the Sauerthal-Schichten, Bornich-Schichten and Kaub-Schichten. These are 408–400 Mya old, making them part of the Latest Pragian to Early Emsian stages of the Devonian.
History of mining
The Hunsrück slate was a source for Rhenish slate over several centuries. Archaeological finds in West Germany show that the slate was used in Roman times. The first documented case of mining in this area dates from the 14th century.[2] The production continued with the Industrial Revolution at the end of the 1700s, but in 1846–49, the industry fell into crisis, resulting in poverty and misery in the mining areas.[3]
The economic upturn after the Franco-Prussian War of 1870-71 resulted in a renewed increase in slate production, where companies used more extensive pits. Production continued until the 1960s, when the competition from cheaper synthetic or imported slate resulted in production decline. Only a single pit in the Bundenbach region was worked in the 1990s. Since 1999, slate imports from Spain, Portugal, Argentina and China caused the abandonment of local mining.
Mining of Hunsrück slate was important for the discovery of fossils. Although not rare, fossils can only be found through extensive mining of slate. Many of the fine fossils exhibited in museums today were originally found by the slate miners. The first scientific publication on these fossils comes from Ferdinand von Roemer (1862),[4] who described starfish and crinoid fossils from Bundenbach. German paleontologists such as R. Opitz, F. Broili, R. Judge, and W. M. Lehmann studied many fossils between 1920 and 1959. Lehmann's death in 1959 and the decline of the slate industry caused a decline in fossil research.
In 1970, Wilhelm Stürmer, a chemical physicist and radiologist at Siemens, developed a new method to examine the Hunsrück slate fossils using medium energy X-rays of 25-40 keV. He created high-resolution movies and stereoscopic images of unopened slates, which showed complex details of soft tissues that cannot be made visible with conventional methods. In the 1990s, Christoph Bartels and Günther Brassel have continued this work.[2]
Paleontology
The various fossil localities are quarries located mostly south of the River Mosel and west of the Rhine in western Germany. The biota of the Hunsrück Slate are commonly called "Bundenbach fossils" after the nearby German community of Bundenbach. More formally, the Hunsruck Slate is properly designated as a Konservat Lagerstätte due to the many fossils that exhibit preservation of soft tissues.[5]
Preservation and taphonomy
Hunsrück is one of the few marine Devonian Lagerstätte having soft tissue preservation, and in many cases fossils are coated by a pyritic surface layer. Preservation of soft tissues as fossils normally requires rapid burial in an anoxic (i.e., with little or no oxygen) sedimentary layer where the decomposition of the organic matter is significantly slowed. The pyritization found in Bundenbach fossils facilitated preservation and enhanced the inherent beauty of the fossils.[6]
Pyritization is rare in the fossil record, and is believed to require not only rapid burial, but both burial in sediments low in organic matter, and high in concentrations of sulfur and iron. Such pyritization is also prevalent in the lower Cambrian fossils from the Maotianshan shales of Chengjiang, China, the oldest Konservat Lagerstätte of Cambrian time.[7]
The best localities for exceptionally preserved fossils are in the communities of Bundenbach and Gemünden. The slates were widely quarried in the past, mainly for roofing tiles from small pits, of which over 600 are known. Today, only a single quarry remains open in the main fossiliferous region of Bundenbach. There are also areas of the Hunsrück Slates where fossils are neither well preserved, nor pyritized, indicating that there also existed environments with shallow and fully oxygenated water.
Diversity of fauna
More than 260 animal species have been described from the Hunsrück Slate. The deposits occur in a strip some 15 km wide and 150 km long running from northwest to southeast. In the main depositional basins of Kaub, Bundenbach, and Gemünden, echinoderms are concentrated in the southwestern area around Bundenbach, with brachiopods predominating in the northeast. The presence of corals and trilobites with well-developed eyes and the rarity of plant fossils from the central basin areas suggest a shallow-water environment. Other animal fossils include sponges, corals, brachiopods, cephalopods, ctenophores,[2] cnidarians, gastropods, and worm trace fossils.[8] Trilobites and echinoderms are relatively abundant in some horizons. Crinoids and starfish are the predominant representatives of the echinoderms, although holothurians (sea cucumbers) are also represented. More than 60 species of crinoids are described from the Hunsrück Slate.
Many types of fishes have been described from the Hunsruck slate. Several genera of placoderm armoured fish have been recorded, including some preserved in three dimensions.[9] Agnathan jawless fishes are the most commonly preserved vertebrates, particularly the flattened Drepanaspis, notable for its upwards-facing mouth, and the streamlined Pteraspis. Spines from acanthodii[10] spiny sharks and a single sarcopterygian lobe-fin specimen are also known.[11][8]
Paleobiota
Color key
|
Notes Uncertain or tentative taxa are in small text; |
Arthropods
Crustaceans
| Crustaceans of the Hunsrück Slate. | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Nahecaris | A phyllocarid crustacean |  | ||
| Oryctocaris | A phyllocarid crustacean | |||
| Ceratiocaris[12] | A phyllocarid crustacean | |||
| Heroldina | A phyllocarid crustacean | |||
| Hohensteiniella[13] | Early crustacean | |||
Eurypterids
| Eurypterids of the Hunsrück Slate. | ||||
|---|---|---|---|---|
| Jaekelopterus | A giant eurypterid, probably allochthonous[14] |  | ||
|
Probably misidentified remains of trilobites[14] |
||||
Marrellomorphs
| Marrellomorphs of the Hunsrück Slate. | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Mimetaster | 123 specimens | A marrellomorph, most common non-trilobite arthropod |  | |
| Vachonisia | A marrellomorph |  | ||
Sea spiders
| Sea spiders of the Hunsrück Slate. | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Flagellopantopus | 1 specimen | A sea spider |  | |
| Palaeopantopus | 3 specimens | A sea spider |  | |
| Palaeoisopus | Over 80 specimens | A sea spider, most common sea spider. |  | |
| Palaeothea | 1 specimen (unlocatable) | A sea spider, potential nomen dubium |  | |
| Pentapantopus | 3 specimens | A sea spider |  | |
Trilobites
| Trilobites of the Hunsrück Slate. | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Chotecops | A trilobite, most common trilobite and arthropod |  | ||
| Odontochile | A trilobite |  | ||
| Cornuproetus | A trilobite | |||
| Parahomalonotus | A trilobite | |||
| Wenndorfia | A trilobite | |||
| Rhenops | A trilobite | |||
| "Asteropyge" | A trilobite | |||
Other Arthropods
| Other arthropods of the Hunsrück Slate. | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Bundenbachiellus | An arthropod of uncertain affinities |  | ||
| Captopodus | Possible early mandibulate related to Acheronauta[15] | |||
| Cambronatus | An arthropod of uncertain affinities | |||
| Magnoculocaris | An arthropod of uncertain affinities, originally named as Magnoculus[16] | |||
| Cheloniellon | A cheloniellid |  | ||
| Palaeoscorpius | A primitive scorpion |  | ||
| Schinderhannes | 1 Specimen | A radiodont |  | |
| Weinbergina | A chelicerate |  | ||
| Wingertshellicus | An arthropod of uncertain affinities |  | ||
Echinoderms
Annelids
| Annelids | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Bundenbachochaeta | A polychaete | |||
| Hunsrueckochaeta | A polychaete | |||
| Ewaldips | A polychaete | |||
| Crocancistrius | A polychaete | |||
| Scopyrites | A polychaete | |||
| Lepidocoleus | A machaeridian | |||
Molluscs
| Annelids | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Anetoceras | An ammonoid cephalopod |  | ||
| Ivoites[17] | An ammonoid cephalopod | |||
| Mimagoniatites | An ammonoid cephalopod | |||
| Mimosphinctes | An ammonoid cephalopod | |||
| Chebbites | An ammonoid cephalopod | |||
| Praecardium | A bivalve | |||
| Panenka | A bivalve | |||
| Ctenodonta | A bivalve | |||
| Cypricardella | A bivalve | |||
| Ctenodonta | A bivalve | |||
| ?Puella | A bivalve | |||
| Palaeozygopleura | A gastropod | |||
| Bembexia | A gastropod | |||
| Loxonema | A gastropod | |||
| Platyceras | A gastropod | |||
Brachiopods
| Brachiopods | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Arduspirifer | A spiriferid brachiopod | |||
| Euryspirifer | ||||
| Brachyspirifer | ||||
| Orbiculoidea | ||||
| Anoplotheca | ||||
| Atrypa | ||||
| Chonetes | ||||
| ?Cryptonella | ||||
| Leptostrophia | ||||
| Loreleiella | ||||
| Meganteris | ||||
| Oligoptycherhynchus | ||||
| Platyorthis | ||||
| Plebejochonetes | ||||
| Tropidoleptus | ||||
Vertebrates
| Vertebrates | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Stuertzaspis | A placoderm | |||
| Tityosteus | A placoderm | |||
| Stensioella | A placoderm |  | ||
| Lunaspis | A placoderm |  | ||
| Nessariostoma | A placoderm |  | ||
| Drepanaspis | A pteraspidomorph jawless fish |  | ||
| Rhinopteraspis | A pteraspidomorph jawless fish |  | ||
| Paraplesiobatis | A placoderm |  | ||
| Pseudopetalichthys | A placoderm |  | ||
| Gemuendina | A placoderm |  | ||
| Westollrhynchus | A lungfish | |||
| Machaeracanthus | Acanthodian | |||
Other animals
| Other animals | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Archaeocydippida | A Ctenophore |  | ||
| Paleoctenophora | A ctenophore |  | ||
| Volgerophyllum[18] | A rugose coral | |||
| "Zaphrentis" | A rugose coral | |||
| Favosites | A tabulate coral | |||
| Pleurodictyum | A tabulate coral | |||
| Palaeonectris | A hydrozoan | |||
| Plectodiscus | A hydrozoan | |||
| Hederella | A hederellid | |||
| Conularia | A conulariid | |||
| Sphenothallus | ||||
| Viriatellina | A tentaculitoid | |||
| Nowakia | ||||
| Styliolina | ||||
| Tentaculites | ||||
| "Protospongia" rhenana | A sponge | |||
| Retifungus | ||||
| Olkenbachia | ||||
| Clionolithes | ||||
Ichnotaxon
| Ichnotaxon | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
|
Worm-like ichnotaxon |
||||
|
Ichnotaxon |
||||
|
Ichnotaxon |
||||
|
Ichnotaxon, including fossils originally misidentified as Edestus[19] |
||||
|
Ichnotaxon |
||||
|
Ichnotaxon |
||||
Plants
| Flora | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Drepanophycales indeterminate[20] | A large lycophyte, washed into the basin from terrestrial sources, similar to Drepanophycus | |||
| Psilophyton | ||||
| Trimerophyton | ||||
| Receptaculites | ||||
| Maucheria | ||||
Fungi
| Fungi | ||||
|---|---|---|---|---|
| Genus | Abundance | Notes | Images | |
| Prototaxites | A fungus that grew in thick, tree-like trunks, washed in from terrestrial sources | |||
See also
- List of fossil sites (with link directory)
References
- ^ Mittmeyer, H.G. (1980). "Zur Geologie des Hunsrückschiefers". Kl. Senckenberg R. (in German). 11. Frankfurt am Main: 26–33.
{{cite journal}}: Cite journal requires|journal=(help) - ^ a b c Bartels, Christoph (2009). The fossils of the Hunsruck slate: Marine life in the Devonian (2nd ed.). Cambridge, UK: Cambridge University Press. pp. 86–88. ISBN 978-0-521-44190-2.
- ^ Wilkin, Jack T. R. (March 2023). "The Hunsrück Slate Konservat‐Lagerstätte: The Hunsrück Slate Konservat‐Lagerstätte". Geology Today. 39 (2): 72–76. doi:10.1111/gto.12426. ISSN 0266-6979.
- ^ von Roemer, Ferdinand (1862). "Neue Asteriden und Crinoiden aus dem devonischen Dachschiefer von Bundenbach bei Birkenfeld". Palaeontographica (in German). 9: 143–52.
- ^ Wilkin, Jack T. R. (March 2023). "The Hunsrück Slate Konservat‐Lagerstätte: The Hunsrück Slate Konservat‐Lagerstätte". Geology Today. 39 (2): 72–76. doi:10.1111/gto.12426. ISSN 0266-6979.
- ^ Wilkin, Jack T. R. (March 2023). "The Hunsrück Slate Konservat‐Lagerstätte: The Hunsrück Slate Konservat‐Lagerstätte". Geology Today. 39 (2): 72–76. doi:10.1111/gto.12426. ISSN 0266-6979.
- ^ Butterfield, Nicholas J. (2003). "Exceptional Fossil Preservation and the Cambrian Explosion". Integrative and Comparative Biology. 43 (1): 166–177. doi:10.1093/icb/43.1.166. PMID 21680421.
- ^ a b "Bundenbach - Lower Devonian Hunsrück Slate of Germany". Fossilmuseum.net. Retrieved 2010-11-27.
- ^ Brazeau; Friedman; Jerve; Atwood (2017). "A three-dimensional placoderm (stem-group gnathostome) pharyngeal skeleton and its implications for primitive gnathostome pharyngeal architecture". Journal of Morphology. 278 (9): 1220–1228. doi:10.1002/jmor.20706. PMC 5575467. PMID 28543631.
- ^ Südkamp, W.H.; Burrow, C.J. (2007). "The acanthodianMachaeracanthus from the Lower Devonian Hunsrück Slate of the Hunsrück region (Germany)". Paläontol Z. 81: 97–104. doi:10.1007/BF02988383. S2CID 129936048.
- ^ Selden, Paul; Nudds, John (2012). Evolution of Fossil Ecosystems. Elsevier Science. p. 68.
- ^ Poschmann, Markus; Bergmann, Alexandra; Kühl, Gabriele (2018). Zhang, Xi-Guang (ed.). "Appendages, functional morphology and possible sexual dimorphism in a new ceratiocaridid (Malacostraca, Phyllocarida) from the Early Devonian Hunsrück Slate (south-western Germany)". Papers in Palaeontology. 4 (2): 277–292. doi:10.1002/spp2.1106. S2CID 91024583.
- ^ Haug, Joachim T.; Poschmann, Markus; Hörnig, Marie K.; Lutz, Herbert (2017). Korn, Dieter (ed.). "A crustacean with eumalacostracan affinities from the Early Devonian Hunsrück Slate (SW Germany)". Papers in Palaeontology. 3 (2): 151–159. doi:10.1002/spp2.1070. S2CID 88601890.
- ^ a b Poschmann, Markus; Bergmann, Alexandra; Kühl, Gabriele (2017-06-01). "First record of eurypterids (Chelicerata, Eurypterida) from the Lower Devonian (Lower Emsian) Hunsrück Slate (SW Germany)". PalZ. 91 (2): 163–169. doi:10.1007/s12542-017-0348-6. ISSN 1867-6812. S2CID 132287510.
- ^ Pulsipher, Mikaela A.; Anderson, Evan P.; Wright, Lauren S.; Kluessendorf, Joanne; Mikulic, Donald G.; Schiffbauer, James D. (2022-12-31). "Description of Acheronauta gen. nov., a possible mandibulate from the Silurian Waukesha Lagerstätte, Wisconsin, USA". Journal of Systematic Palaeontology. 20 (1): 1–24. doi:10.1080/14772019.2022.2109216. ISSN 1477-2019. S2CID 252839113.
- ^ Briggs, Derek E. G.; Bartels, Christoph (2002). "Magnoculocaris, A New Name for the Arthropod Magnoculus Briggs and Bartels, 2001 from the Lower Devonian Hunsruck Slate, Germany". Palaeontology. 45 (2): 419. doi:10.1111/1475-4983.00243. ISSN 0031-0239. S2CID 129700543.
- ^ Stilkerich, Julia; Smrecak, Trisha A.; Baets, Kenneth De (2017-06-30). "3D-Analysis of a non-planispiral ammonoid from the Hunsrück Slate: natural or pathological variation?". PeerJ. 5: e3526. doi:10.7717/peerj.3526. ISSN 2167-8359. PMC 5494166. PMID 28674668. S2CID 23907662.
- ^ Südkamp, Wouter H. (2007). "An atypical fauna in the Lower Devonian Hunsrück Slate of Germany". Paläontologische Zeitschrift. 81 (2): 181–204. doi:10.1007/BF02988392. ISSN 0031-0220. S2CID 129466762.
- ^ Itano, Wayne (2020). "FINAL(?) IDENTIFICATION OF THE FALSE EDESTUS FROM THE HUNSRÜCK SLATE: PROTOVIRGULARIA (A TRACE FOSSIL)".
{{cite journal}}: Cite journal requires|journal=(help) - ^ Poschmann, Markus; Gossmann, Rolf; Matsunaga, Kelly K. S.; Tomescu, Alexandru M. F. (2020). "Characterizing the branching architecture of drepanophycalean lycophytes (Lycopsida): an exceptional specimen from the Early Devonian Hunsrück Slate, southwest Germany, and its paleobiological implications". PalZ. 94 (1): 1–16. doi:10.1007/s12542-018-00443-w. ISSN 0031-0220. S2CID 135090137.
External links
Media related to Hunsrück Lagerstätte at Wikimedia Commons