2021 in paleoentomology

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This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

Clade Amphiesmenoptera

Lepidoptera

New Lepidopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acanthocorona^[1]	Gen. et 7 sp. nov	Valid	Mey, Léger & Lien	Cretaceous Cenomanian	Burmese amber	Myanmar	A Lophocoronidae moth. The type species is A. skalskii The genus also includes A. bowangi, A. kuranishii, A. muelleri, A. sattleri, A. spinifera, & A. wichardi.	Acanthocorona skalskii
Agathiphagama^[1]	Gen. et sp. nov	Valid	Mey, Léger & Lien	Cretaceous Cenomanian	Burmese amber	Myanmar	An agathiphagid aglossatan moth. The type species is A. perdita.	Agathiphagama perdita
Phyllocnistis cretacea^[2]	Sp. nov	Valid	Fischer	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Phyllocnistis
Phyllonorycter inopinata^[2]	Sp. nov	Valid	Fischer	Lutetian	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Phyllonorycter

Lepidopteran research

The first known Cretaceous caterpillar armoured with spines is described from the Burmese amber by Haug & Haug (2021).^[3]
Fossil leaf mine produced by a lyonetiid-like leaf-mining moth, representing the oldest record of such moths reported to date, is described from the Upper Cretaceous (Campanian) Kaiparowits Formation (Utah, United States) by Maccracken et al. (2021), who name a new ichnotaxon Leucopteropsa spiralae.^[4]

Trichopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Archaeopolycentra achupomotet^[5]	Sp. nov	Valid	Melnitsky & Ivanov	Late Cretaceous (Santonian)	Kheta Formation (Taymyr amber)	Russia ( Krasnoyarsk Krai)	A member of the family Polycentropodidae.
Archaeopolycentra turvalsia^[5]	Sp. nov	Valid	Melnitsky & Ivanov	Late Cretaceous (Santonian)	Kheta Formation (Taymyr amber)	Russia ( Krasnoyarsk Krai)	A member of the family Polycentropodidae.
Cernotina fossilinova^[6]	Sp. nov	Valid	Wichard & Neumann	Probably early Miocene	Dominican amber	Dominican Republic	A caddisfly, a species of Cernotina.
Electrotrichia rasnitsyni^[7]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Late Eocene	Rovno amber	Ukraine	A member of the family Hydroptilidae.
Folindusia aenigmatosa^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
Folindusia expansa^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
Folindusia inconspicua^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
Folindusia inventa^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
Folindusia robusta^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
Folindusia tortilis^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
Holocentropus tutkaktut^[9]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Eocene (Priabonian)	Rovno amber	Ukraine	A species of Holocentropus.
Palaeopsilotreta kachini^[10]	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber	Myanmar	A member of the family Odontoceridae.
Palaeopsilotreta succini^[10]	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber	Myanmar	A member of the family Odontoceridae.
Plectrocnemia kirmikhia^[11]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Eocene (Priabonian)	Rovno amber	Ukraine	A species of Plectrocnemia.
Psilotreta fossilis^[10]	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber	Myanmar	A species of Psilotreta.
Purbimodus khramovi^[12]	Sp. nov	Valid	Sukatsheva & Aristov	Early Cretaceous		Russia	A member of the family Vitimotauliidae.
Terrindusia nova^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
Wormaldia denticulata^[13]	Sp. nov	In press	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A caddisfly, a species of Wormaldia.
Wormaldia diplobifurca^[13]	Sp. nov	In press	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A caddisfly, a species of Wormaldia.

Clade Antliophora

Dipterans

New Dipteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acartophthalmites crassipes^[14]	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea belonging to the new family Clusiomitidae.
Acartophthalmites luridus^[14]	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea belonging to the new family Clusiomitidae.
Acartophthalmites rugosus^[14]	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea belonging to the new family Clusiomitidae.
Aleksiana^[15]	Gen. et sp. et comb. nov	Valid	Krzemiński et al.	Cretaceous	Burmese amber	Myanmar	A member of the crane fly family Limoniidae. The type species is A. rasnitsyni; genus also includes "Antocha" lapra Podenas & Poinar (2009).
Allarete patriciae^[16]	Sp. nov	Valid	Nel	Early Eocene	Oise amber	France	A member of the family Cecidomyiidae.
Allodia paleoafricana^[17]	Sp. nov	Valid	Bouju et al.	Early Miocene	Ethiopian amber	Ethiopia	A species of Allodia.
Apocephalus chiapanecus^[18]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Mexican amber	Mexico	A species of Apocephalus.
Apocephalus dominicanus^[18]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Dominican amber	Dominican Republic	A species of Apocephalus.
Apocephalus miocenus^[18]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene			A species of Apocephalus.
Araripogon pulchrus^[19]	Sp. nov	In press	Lamas, Sampronha & Ribeiro	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Asilidae.
Bashkonia^[20]	Gen. et sp. nov	Valid	Lukashevich	Middle Triassic	Röt Formation	Germany	A member of the family Nadipteridae. Genus includes new species B. franconica.
Bibio castaneipennis^[21]	Sp. nov	Valid	Skartveit & Wedmann	Oligocene		Germany	A species of Bibio.
Bibio succineus^[22]	Sp. nov	Valid	Skartveit	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Bibio.
Burmahesperinus^[23]	Gen. et 3 sp. nov	Valid	Ševčík et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Bibionidae. The type species is B. antennatus; genus also includes B. conicus and B. pedicellatus.
Burmasymmerus^[24]	Gen. et 2 sp. nov	Valid	Ševčík, Hippa & Burdíková	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Ditomyiidae. The type species is B. korneliae; genus also includes B. wieslawi.
Burmatricha^[24]	Gen. et sp. nov	Valid	Ševčík, Hippa & Burdíková	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Sciaroidea of uncertain phylogenetic placement. The type species is B. mesozoica.
Cascomixticus^[25]	Gen. et sp. nov	Valid	Poinar & Vega	Cretaceous (Albian/Cenomanian)	Burmese amber	Myanmar	A member of the family Apsilocephalidae. The type species is C. tubuliferous.
Clusiomites^[14]	Fam. et gen. et comb. et sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea, the type genus of the new family Clusiomitidae. The type species is "Acartophthalmites" clusioides Roháček (2016); genus also includes new species C. ornatus.
Crenoptychoptera gaoi^[26]	Sp. nov	Valid	Liu & Huang	Middle–Late Jurassic	Jiyuan Basin	China	A member of the family Ptychopteridae.
Cretobibio rasnitsyni^[27]	Sp. nov	Valid	Lukashevich, Amorim & Ribeiro	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Bibionidae.
Cretolimonia dayana^[28]	Sp. nov		Kopeć in Kopeć et al.	Jurassic/Cretaceous boundary	Glushkovo Formation	Russia ( Zabaykalsky Krai)	A member of the subfamily Architipulinae.
Cretolimonia lukashevichae^[28]	Sp. nov		Kopeć & Krzemiński in Kopeć et al.	Jurassic/Cretaceous boundary	Dain Formation	Russia ( Zabaykalsky Krai)	A member of the subfamily Architipulinae.
Cretolimonia mikolajczyki^[28]	Sp. nov		Kopeć, Krzemiński, Soszyńska-Maj in Kopeć et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the subfamily Architipulinae.
Cretolimonia pseudojurassica^[28]	Sp. nov		Krzemiński in Kopeć et al.	Jurassic/Cretaceous boundary	Dain Formation	Russia ( Zabaykalsky Krai)	A member of the subfamily Architipulinae.
Cretothereva^[29]	Gen. et sp. nov	In press	Carmo, Lamas & Ribeiro	Early Cretaceous	Crato Formation	Brazil	A member of the family Therevidae. Genus includes new species C. antiqua.
Cretpenthetria^[30]	Gen. et sp. nov	Valid	Li, Zhang & Xiao	Cretaceous	Burmese amber	Myanmar	A member of the family Bibionidae. Genus includes new species C. burmensis.
Culex ekaterinae^[31]	Sp. nov	Valid	Giłka, Harbach & Perkovsky	Eocene	Rovno amber	Ukraine	A mosquito, a species of Culex.
Culicoides paleopestis^[32]	Sp. nov	In press	Peñalver et al.	Eocene	Anglesea amber	Australia	A species of Culicoides.
Cyttaromyia freiwaldi^[33]	Sp. nov	Valid	Kania-Kłosok et al.	Eocene	Green River Formation	United States	A member of the family Cylindrotomidae.
Cyttaromyia gelhausi^[33]	Sp. nov	Valid	Kania-Kłosok et al.	Eocene	Green River Formation	United States	A member of the family Cylindrotomidae.
Decessia^[34]	Gen. et sp. nov	Valid	Krzemiński et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the crane fly family Limoniidae. The type species is D. podenasi.
Elephantomyia christelae^[35]	Sp. nov		Kania-Kłosok & Krzemiński	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Elephantomyia.
Elephantomyia prima^[35]	Sp. nov		Kania-Kłosok & Krzemiński	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Elephantomyia.
Forcipomyia rasnitsyni^[36]	Sp. nov	Valid	Szadziewski, Sontag & Pankowski	Miocene	Ethiopian amber	Ethiopia	A species of Forcipomyia.
Helius hispanicus^[37]	Sp. nov		Kania-Kłosok, Krzemiński & Arillo	Early Cretaceous (Albian)	Utrillas Group (Álava amber)	Spain	A species of Helius.
Helius turolensis^[37]	Sp. nov		Kania-Kłosok, Krzemiński & Arillo	Early Cretaceous (Albian)	Utrillas Group (San Just amber)	Spain	A species of Helius.
Hirmoneura messelense^[38]	Sp. nov	Valid	Wedmann et al.	Eocene (Lutetian)	Messel Formation	Germany	A member of the family Nemestrinidae.
Lebambromyia sacculifera^[39]	Sp. nov	Valid	Badano, Zhang, Yeates & Cerretti in Badano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Phoroidea.
Leptotarsus andradei^[40]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
Leptotarsus gelhausi^[40]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
Leptotarsus ivoneae^[40]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
Leptotarsus lemeae^[40]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
Mesoacentron^[41]	Gen. et sp. nov	Valid	Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.	Late Cretaceous (Santonian)	Kheta Formation (Taimyr amber)	Russia	A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is M. kaluginae.
Meunierohelea anglesensis^[32]	Sp. nov	In press	Peñalver et al.	Eocene	Anglesea amber	Australia	A member of the family Ceratopogonidae.
Minyohelea nexuosa^[42]	Sp. nov	In press	Pielowska-Ceranowska, Azar & Szwedo	Early Cretaceous (Barremian)	Lebanese amber	Lebanon	A member of the family Ceratopogonidae.
Myriophora asiatica^[18]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Zhangpu amber	China	A species of Myriophora.
Neoarisemus groehni^[43]	Sp. nov	Valid	Wagner	Eocene		Ukraine	A member of the family Psychodidae.
Orimarguloides^[44]	Gen. et sp. nov	Valid	Men & Podenas	Cretaceous	Burmese amber	Myanmar	A member of the crane fly family Limoniidae. Genus includes new species O. simplex.
Palaeocentron^[41]	Gen. et sp. nov	Valid	Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.	Cretaceous (probably Albian–Cenomanian)	Burmese amber	Myanmar	A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is P. krzeminskii.
Palaeoglaesum pilosus^[45]	Sp. nov	Valid	Skibińska, Krzemiński & Zhang in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Palaeoglaesum stebneri^[45]	Sp. nov	Valid	Skibińska & Krzemiński in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Palaeoglaesum teres^[45]	Sp. nov	Valid	Skibińska & Albrycht in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Paleoplatyura agnieszkae^[46]	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber	Myanmar	A species of Paleoplatyura.
Paleoplatyura magnifica^[46]	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber	Myanmar	A species of Paleoplatyura.
Paleoplatyura miae^[46]	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber	Myanmar	A species of Paleoplatyura.
Paleotelmatoscopus^[47]	Gen. et comb. et sp. nov	Valid	Curler & Skibińska	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Psychodidae. The type species is "Pericoma" formosa Meunier (1905); genus also includes new species P. madrizi.
Polymera alexanderi^[48]	Sp. nov	Valid	Krzemiński et al.	Miocene	Dominican amber	Dominican Republic	A species of Polymera.
Protopenthetria^[49]	Gen. et sp. nov	Valid	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Bibionidae. Genus includes new species P. skartveiti.
Rhabdomastix asiatica^[50]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
Rhabdomastix cretacica^[50]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
Rhabdomastix krzeminskae^[50]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
Rhabdomastix myanmae^[50]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
Rhipidoxylomyia rasnitsyni^[16]	Sp. nov	Valid	Nel	Early Eocene	Oise amber	France	A member of the family Cecidomyiidae.
Stempellinella gilkai^[51]	Sp. nov	Valid	Zakrzewska & Jankowska	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Stempellinella.
Trichoneura xavieri^[52]	Sp. nov	Valid	Kania-Kłosok et al.	Early Cretaceous (Albian)	Utrillas Group	Spain	A species of Trichoneura.
Zhangsolva burmensis^[53]	Sp. nov	In press	Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Zhangsolvidae.

Dipteran research

Baranov et al. (2021) document a high diversity of fly larvae from the Eocene Baltic amber, preserving taxonomically, morphologically, and ecologically important characters, and evaluate the implications of this finding for the knowledge of the functional ecology of the Eocene fly larvae.^[54]

Mecopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Conicholcorpa longa^[55]	Sp. nov		Zhang, Shih & Ren in Zhang et al.	Middle Jurassic	Jiulongshan Formation	China	A scorpionfly belonging to the family Holcorpidae.
Glanomerope^[56]	Gen. et sp. nov	Valid	Poschmann & Nel	Permian	Meisenheim Formation	Germany	A scorpionfly belonging to the family Protomeropidae. Genus includes new species G. virgoferroa.
Longiphlebia incompleta^[57]	Sp. nov	In press	Lian, Cai & Huang	Late Jurassic	Tiaojishan Formation	China	A scorpionfly belonging to the family Orthophlebiidae.
Mesopsyche jinsuoguanensis^[58]	Sp. nov	Valid	Lian, Cai & Huang	Middle Triassic	Tongchuan Formation	China	A member of the family Mesopsychidae. Originally described as a species of Mesopsyche; transferred by Lian et al. (2024) to the genus Turbidapsyche.^[59]
Mesopsyche liaoi^[58]	Sp. nov	Valid	Lian, Cai & Huang	Late Triassic	Huangshanjie Formation	China	A member of the family Mesopsychidae. Originally described as a species of Mesopsyche; transferred by Lian et al. (2024) to the genus Turbidapsyche.^[59]

Clade Archaeorthoptera

†Caloneurodea

Name	Novelty	Status	Authors	Age	Country	Notes
Aviobiella^[60]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)	France	A member of Archaeorthoptera belonging to the order Caloneurodea. Genus includes new species A. garroustei.
Oudardgramma^[61]	Gen. et sp. nov		Nel & Roques	Carboniferous (Bashkirian)	France	The type species is O. bruayensis.
Paraspiculum^[62]	Gen. et sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)	Germany	A member of Archaeorthoptera belonging to the group Caloneurodea. Genus includes new species P. brauckmannii.

†Cnemidolestodea

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Avionxixia^[63]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)		France	A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is A. gui.
Ctenoptilus frequens^[64]	Sp. nov	Valid	Chen et al.	Carboniferous (Bashkirian to Moscovian)	Yanghugou Formation	China	A member of Archaeorthoptera belonging to the family Ctenoptilidae.
Palatinarkema^[65]	Gen. et sp. nov	Valid	Nel & Poschmann	Early Permian (probably Asselian–?Sakmarian)	Meisenheim Formation	Germany	A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is P. prokopi.
Piesbergopterum^[62]	Gen. et 2 sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)		France Germany	A member of Archaeorthoptera belonging to the group Cnemidolestodea. Genus includes new species P. punctatum Dvořák et al. and P. avionensis Nel & Roques.^[66]

Orthopterans

New Orthopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Amberotridactylus^[67]	Gen. et sp. nov	In press	Du, Xu & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Tridactylidae. Genus includes new species A. cheni.
Archaboilus polyneurus^[68]	Sp. nov	Valid	Gu, Yue & Ren in Gu et al.	Middle Jurassic (Bathonian–Callovian)	Jiulongshan Formation	China	An ensiferan belonging to the group Hagloidea and the family Haglidae.
Caelielca^[69]	Gen. et sp. nov	In press	Uchida	Cretaceous	Burmese amber	Myanmar	A member of the family Elcanidae. Genus includes new species C. spinocrus.
Cascogryllus setosus^[67]	Sp. nov	In press	Du, Xu & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Tridactylidae.
Cratoelcana rasnitsyni^[70]	Sp. nov	In press	Nel & Jouault	Early Cretaceous	Crato Formation	Brazil	A member of the family Elcanidae.
Hukawnelca^[69]	Gen. et sp. nov	In press	Uchida	Cretaceous	Burmese amber	Myanmar	A member of the family Elcanidae. Genus includes new species H. gracile.
Panorpidium spica^[71]	Sp. nov	In press	Kim et al.	Early Cretaceous (Albian)	Jinju Formation	South Korea	An orthopteran belonging to the family Elcanidae.
Permohagla^[72]	Gen. et sp. nov	Valid	Gorochov	Late Permian		Russia ( Kemerovo Oblast)	A member of Hagloidea. Genus includes new species P. tomica.
Phyllotridactylus^[73]	Gen. et sp. nov	In press	Xu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Tridactylidae. Genus includes new species P. wangi.
Pseudoacrida sennlaubi^[70]	Sp. nov	In press	Nel & Jouault	Early Cretaceous	Crato Formation	Brazil	A member of the family Locustopsidae.

Orthopteran research

Revision of the fossil record of the family Trigonidiidae is published by Desutter-Grandcolas et al. (2021).^[74]

†Titanoptera

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Theiatitan^[75]	Fam. et gen. et sp. nov	Valid	Schubnel, Roques & Nel in Schubnel et al.	Carboniferous (Moscovian)		France	A member of Titanoptera belonging to the new family Theiatitanidae. The type species is T. azari.	Theiatitan azari

Other archaeorthopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Archaeogerarus^[63]	Fam. et gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)		France	A member of Archaeorthoptera belonging to the new family Archaeogeraridae. The type species is A. schubneli.
Avionugonioneura^[76]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)		France	A member of Archaeorthoptera possibly belonging to the family Nugonioneuridae. Genus includes new species A. jouaulti.
Contracladus^[62]	Gen. et sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)		Germany	A member of Archaeorthoptera of uncertain phylogenetic placement. Genus includes new species C. impar.
Duquesnia^[77]	Gen. et sp. nov	Valid	Nel & Roques in Nel, Roques & Duquesne	Carboniferous (Moscovian)		France	A member of Archaeorthoptera. Genus includes new species D. gallica.
Penanoptera^[78]	Gen. et sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin	Germany	A member of Archaeorthoptera. Genus includes new species P. manseri.

Clade Coleopterida

Coleoptera

Adephaga

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Antephilorhizus^[79]	Gen. et sp. nov	Valid	Kirichenko-Babko, Perkovsky & Vasilenko	Eocene	Rovno amber	Ukraine	A ground beetle belonging to the tribe Lebiini. Genus includes new species A. pripiatiensis.
Balticeler^[80]	Gen. et sp. nov	Valid	Schmidt & Maddison in Schmidt, Scholz & Maddison	Eocene	Baltic amber	Europe (Baltic Sea region)	A ground beetle belonging to the subfamily Trechinae. The type species is B. kerneggeri.
Calosoma kimi^[81]	Sp. nov	Valid	Lee & Nam	Early Miocene	Geumgwangdong Formation	South Korea	A species of Calosoma.

Archostemata

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Jurodes shef^[82]	Sp. nov	Valid	Yan, Strelnikova & Cai	Early Cretaceous (Berriasian-Barremian)	Gusinoe Ozero Group	Russia ( Buryatia)	A Jurodid beetle.
Kirejtomma^[83]	Gen. et comb. nov	Valid	Li & Cai	Cretaceous	Burmese amber	Myanmar	An Ommatid beetle The type species is "Clessidromma" zengi (2020).	Kirejtomma zengi
Limnomma^[84]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Middle Jurassic	Haifanggou Formation	China	An Ommatid beetle. The type species is L. daohugouense.	Limnomma daohugouense
Omma axsmithi^[85]	Sp. nov	In press	Jarzembowski, Zheng & Zhao	Cretaceous	Burmese amber	Myanmar	An Omma species Ommatid beetle.
Omma forte^[86]	Sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	An Omma species Ommatid beetle.
Paraodontomma leptocristatum^[87]	Sp. nov	Valid	Li, Yamamoto & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	An Ommatid beetle.	Paraodontomma leptocristatum
Permocatinus^[88]	Gen. et sp. nov	Valid	Ponomarenko	Late Permian		Russia	A Coleocatiniid beetle. The type species is P. tomiensis.
Tomiaplus^[88]	Gen. et comb. et sp. nov	Valid	Ponomarenko	Late Permian		Russia	A Triaplid beetle. The type species is "Triaplus" sibiricus (2013), as well as new species T. minimus.
Tunguskagyrus уani^[88]	Sp. nov	Valid	Ponomarenko	Late Permian		Russia	A Triaplid beetle.
Zygadenia alexrasnitsyni^[89]	Sp. nov	Valid	Strelnikova & Yan	Early Cretaceous	Zaza Formation	Russia	An Ommatid beetle.

Myxophaga

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Triamyxa^[90]	Fam., Gen. et sp. nov	Valid	Qvarnström et al.	Late Triassic (Carnian)	Keuper succession (Drawno Beds)	Poland	A Triamyxid beetle in the suborder Myxophaga. The type species is T. coprolithica.	Triamyxa coprolithica

Polyphaga

Bostrichiformia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Archaenosodendron^[91]	Gen. comb. et 3 sp. nov	Valid	Li & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	A nosodendrid beetle. The type species is "Nosodendron" cretaceum; genus also includes new species A. angulare, A. explanatum and A. remotidens.
Cretomegatoma^[92]	Gen. et comb. nov	Valid	Háva	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A carpet beetle. The type species is "Megatoma" atypica (2017).
Mesonosa^[93]	Gen. et sp. nov	Valid	Tihelka et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A wounded-tree beetle. The type species is M. scandens.
Petalium bruteno^[94]	Sp. nov	Valid	Alekseev & Bukejs	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A Petalium species death watch beetle.
Petalium widewuto^[94]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Petalium species death watch beetle.
Trogoderma ainu^[95]	Sp. nov	Valid	Perkovsky, Háva & Zaitsev	Middle Eocene	Sakhalin amber	Russia	A Trogoderma species carpet beetle.
Tuberphradonoma^[92]	Gen. et sp. nov	Valid	Háva	Late Cretaceous Cenomanian)	Burmese amber	Myanmar	A carpet beetle. The type species is T. burmitica.
Xyletinus lobanovi^[96]	Sp. nov	Valid	Bukejs, Alekseev & Háva	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Xyletinusspecies death watch beetle.

Cucujiformia

New Cucujiformian taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Aberrokorynetes oceanojubilaei^[97]	Sp. nov	In press	Kolibáč et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Korynetinae checkered beetle.
Aliattalus^[98]	Gen. et sp. nov	Valid	Tshernyshev	Late Eocene	Baltic amber	Europe	A Malachiinae malachite beetle. The type species is A. intercalaris.
Archaeosciaphilus gyrommatus^[99]	Sp. nov	In press	Legalov & Poinar	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Polydrusini Entiminae broad nosed weevil.
Arkagalarhinus^[100]	Gen. et sp. nov	In press	Legalov	Late Cretaceous (Santonian-Campanian)	Arkagala Formation	Russia	A Chilecarinae ithycerid weevil. The type species is A. zherikhini.
Asiomira dubrovinae^[101]	Sp. nov	Valid	Nabozhenko & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	An Alleculinae darkling beetle.
Aulacoscelis santanensis^[102]	Sp. nov	In press	Legalov	Early Cretaceous (Aptian-Albian)	Santana Group	Brazil	An orsodacnid leaf beetle.
Baltocar sontagae^[103]	Sp. nov	Valid	Bukejs & Legalov	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Sayrevilleinae tooth-nosed snout weevil.
Baltosidis^[104]	Gen. et 3 sp. nov	Valid	Szawaryn	Eocene	Baltic amber	Europe	A Microweiseini Microweiseinae lady beetle. The type species is B. damzeni; genus also includes B. damgaardi and B. szadziewskii.
Bronchotibia^[105]	Gen. et sp. nov	In press	Poinar & Legalov	Miocene (Burdigalian)	Dominican amber	Dominican Republic	A Lymantini Molytinae weevil. The type species is B. adunatus.
Cathartosilvanus perkovskyi^[106]	Sp. nov	Valid	Alekseev & Bukejs	Late Eocene	Rovno amber	Ukraine	A silvanid beetle.
Ceutorhynchus zerovae^[107]	Sp. nov	Valid	Legalov et al.	Late Eocene	Rovno amber	Ukraine	A weevil.
Coslonatus^[108]	Gen. et sp. nov	Valid	Li & Cai	Cenomanian	Burmese amber	Myanmar	A Colydiinae? Zopheridae tenebrionoid beetle. The type species is C. rasnitsyni.
Cretaidgia^[109]	Gen. et sp. nov	In press	Zhao, Liu & Yu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A prionocerid beetle. The type species is C. burmensis.
Diapus ethiopicus^[110]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Ethiopian amber	Ethiopia	A Platypodinae pinhole borer ambrosia weevil.
Diapus resinae^[110]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Zhangpu amber	China	A Platypodinae pinhole borer ambrosia weevil.
Dorytomus mikhailovi^[111]	Sp. nov	Valid	Legalov, Nazarenko & Perkovsky	Late Eocene	Rovno amber	Ukraine	A species of Dorytomus Curculioninae weevil.
Electroxylita^[112]	Gen. et sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle. The type species is E. chronographica.
Eurocrossotus^[113]	Gen. et sp. nov	Valid	Vitali	Eocene	Baltic amber	Europe (Baltic Sea region)	A longhorn beetle belonging to the tribe Crossotini. The type species is E. alekseevi.
Glaesotropis rohdendorfi^[114]	Sp. nov	Valid	Legalov, Nazarenko & Perkovsky	Eocene (Priabonian)	Rovno amber	Ukraine	A zygaenodinin anthribine anthribid fungus weevil.
Glesoconomorphus ekaterinae^[115]	Sp. nov	Valid	Telnov et al.	Late Eocene Priabonian	Rovno amber	Ukraine	A mycterid palm beetle.
Gracilenticrus^[116]	Gen. et sp. nov	Valid	Yu, Kolibáč & Ślipiński in Yu et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A lophocaterid beetle. The type species is G. burmiticus.
Heterhelus buzina^[117]	Sp. nov	Valid	Kupryjanowicz, Lyubarsky & Perkovsky	Eocene	Rovno amber	Ukraine	A Kateretid short-winged flower beetle. Originally described as a species of Heterhelus, but Kirejtshuk, Jenkins Shaw & Smirnov (2023) transferred it to the genus Antirhelus.^[118]
Isomira lobanovi^[119]	Sp. nov	Valid	Nabozhenko & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	An Isomira species claw comb darkling beetle.
Kaszubia^[120]	Gen. et sp. nov	Valid	Legalov, Kupryjanowicz & Perkovsky	Late Eocene Priabonian	Baltic amber	Poland	A cossonine weevil. The type species is K. wanati.
Kulindrobor^[121]	Gen. et 2 sp. nov	Valid	Tihelka et al.	Late Cretaceous Cenomanian	Burmese amber	Myanmar	A tenebrionoid beetle of uncertain phylogenetic placement. Genus includes new species K. enigmaticus and K. magnus.
Lobanoviella^[122]	Gen. et sp. nov	In press	Kirejtshuk & Reid	Eocene	Baltic amber	Europe	A lobanovielline palophagine megalopodid leaf beetle. The type species is L. andreyi.
Madelinia capillata^[112]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
Melanosiagon^[123]	Gen. et sp. nov	Valid	Batelka & Prokop	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A wedge-shaped beetle. The type species is M. serraticornis.
Metaclisa ottoi^[124]	Sp. nov	Valid	Nabozhenko, McKellar & Bukejs	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	AMetaclisa species metaclisine darkling beetle.
Microscapha kugelanni^[112]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
Microtrogossita^[125]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	A Trogossitid bark-gnawing beetle. The type species is M. qizhihaoi.
Monolepta rappsilberi^[126]	Sp. nov	Valid	Bukejs et al.	Late Eocene Priabonian	Bitterfeld amber	Germany	A Monolepta species skeletonizing leaf beetle.
Neolatitergum^[127]	Nom. nov	Valid	Zhang	Jurassic	Jiulongshan Formation	China	A Trogossitid bark-gnawing beetle. A replacement name for "Latitergum" Yu et al. (2014).
Oedemera girulskii^[128]	Sp. nov	Valid	Szawaryn, Sontag & Kubisz	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	An Oedemera species false blister beetle.
Pacyclaxyra^[129]	Gen. et sp. nov	Valid	Tihelka, Huang & Cai	Late Cretaceous Cenomanian	Burmese amber	Myanmar	A sooty mould beetle. The type species is P. azari.
Palaeatalasis^[130]	Gen. et sp. nov	Valid	Legalov	Eocene (Ypresian)	Green River Formation	United States ( Utah)	A Megamerini Sagrinae leaf beetle. The type species is P. monrosi.
Palaeoanoplus^[131]	Trib., Gen., et sp. nov	Valid	Legalov	Late Eocene Priabonian	Baltic amber	Europe	A Palaeoanoplini Curculioninae weevil. The type species is P. horridus.
Paleopegorhinus^[132]	Gen. et sp. nov	Valid	Poinar, Brown & Legalov	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A Chilecarinae new york weevil. The type species is P. micrommatus.
Palaeopeplus^[133]	Gen. et sp. nov	Valid	Powell & Cline	Miocene (Burdigalina)	Dominican amber	Dominican Republic	A Cillaeinae sap beetle. The type species is P. cascus.
Pelretes^[134]^[135]^[136]	Gen. et sp. nov	Valid	Tihelka et al.	Cretaceous	Burmese amber	Myanmar	Originally described as a short-winged flower beetle; subsequently argued to be a sap beetle belonging to the subfamily Apophisandrinae^[137] or a member of the separate family Apophisandridae.^[138] The type species is P. vivificus .
Phloiotrya inmarinata^[112]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A Phloiotrya species false darkling beetle.
Protodasytes^[139]	Gen. et sp. nov	Valid	Tihelka et al.	Late Cretaceous (Cenomanian)	Charentese amber	France	A soft-wing flower beetle. The type species is P. cretaceus.
Protomauroania^[140]	Gen. et 2 sp. nov	Valid	Tshernyshev in Tshernyshev & Shcherbakov	Late Eocene	Baltic amber Rovno amber	Baltic Sea region Ukraine	A Dasytinae malachite beetle. The type species P. chaetophorata Genus also includes P. mikhailovi.^[141]
Protostomopsis^[142]	Gen. et sp. nov	Valid	Bukejs, Ślipiński & Alekseev in Bukejs et al.	Eocene	Baltic amber	Poland Russia ( Kaliningrad Oblast)	A cerylonid beetle. The type species is P. pandema.
Pseudomesauletes lobanovi^[103]	Sp. nov	Valid	Bukejs & Legalov	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Rhynchitinae tooth-nosed snout weevil.
Pulchritudo^[143]	Gen. et sp. nov	Valid	Krell & Vitali	Eocene (Ypresian)	Green River Formation	United States ( Colorado)	A Sagrinae leaf beetle. The type species is P. attenboroughi.
Rhamphophorus^[144]	Trib., gen. et sp. nov	Valid	Poinar & Brown	Cretaceous Cenomanian	Burmese amber	Myanmar	A Rhamphophorini Cimberidinae pine flower weevil. The type species is R. legalovii.
Salignacicola^[123]	Gen. et comb. nov	Valid	Batelka & Prokop	Late Cretaceous (Cenomanian)		France	A wedge shaped beetle. The type species is "Macrosiagon" ebboi (2004).
Symphora glaesonauta^[112]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
Symphora pollocki^[112]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
Teredolaemus primigenius^[145]	Sp. nov	Valid	Alekseev et al.	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A teredid beetle.
Toxorhynchus michalskii^[146]	Sp. nov	Valid	Legalov	Late Eocene Priabonian	Baltic amber	Europe	An Apioninae straight-snouted weevil.
Trematosphindus^[147]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A slime mold beetle. The type species is T. newtoni
Zaiwa^[148]	Gen. et sp. nov	Valid	Lyubarsky et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	A lophocaterid beetle. The type species is Z. pankowskiorum.
Zherikhiniellus^[146]	Gen. et sp. nov	Valid	Legalov	late Eocene Priabonian	Baltic amber	Europe	A Nanophyinae straight-snouted weevil. The type species is Z. rarus.

Cucujiformia research

A piece of Burmese amber preserving the complete series of free-living stages (except of the last larval instar) of a Cretaceous parasitoid beetle Paleoripiphorus is described by Batelka, Engel & Prokop (2021).^[149]

Elateriformia

New Elateriformian taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Amberophytum maculatum^[150]	Sp. nov	Valid	Poinar, Vega & Legalov	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cerophytid rare click beetle.
Anoeuma^[151]	Gen. et sp. nov	Valid	Li, Kundrata & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	An elateroid click beetle of uncertain phylogenetic placement. The type species is A. lawrencei.
Arturmiles^[152]	Gen. et sp. nov	Valid	Fanti	Eocene	Prussian Formation (Baltic amber)	Poland	A soldier beetle belonging to the tribe Cantharini. The type species is A. pankowskiarum.
Balistica vetai^[153]	Sp. nov	Valid	Muona	Eocene	Baltic amber	Europe	A Dirhagini Melasinae false click beetle.
Baltodascillus^[154]	Gen. et sp. nov	Valid	Kundrata et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A dascillid beetle. The type species is B. serraticornis.
Burmagrilus^[155]	Gen. et sp. nov	Valid	Jiang, Song & Wang in Jiang et al.	Cretaceous	Burmese amber	Myanmar	A metallic wood-boring beetle. The type species is B. cretacus.
Burmomiles bilineatimaculatus^[156]	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber	Myanmar	A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.^[157]
Burmomiles laticollis^[156]	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber	Myanmar	A soldier beetle.
Burmomiles oblongoculus^[156]	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber	Myanmar	A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.^[157]
Captopus^[158]	Gen. et sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A throscid beetle. The type species is C. depressiceps.
Chelonarium andabata^[159]	Sp. nov	Valid	Alekseev & Bukejs in Alekseev et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A turtle beetle.
Chelonarium dingansich^[159]	Sp. nov	Valid	Alekseev & Bukejs in Alekseev et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A turtle beetle.
Chelonarium dominicanum^[160]	Sp. nov	Valid	Poinar & Háva	Burdigalian	Dominican amber	Dominican Republic	A turtle beetle.
Coomanius balticus^[153]	Sp. nov	Valid	Muona	Eocene	Baltic amber	Europe	A Nematodini Macraulacinae false click beetle.
Cretocantharis^[161]	Gen. et sp. nov	Valid	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A soldier beetle. The type species is C. veda.
Cretopachyderes^[162]	Gen. et sp. nov	Valid	Kundrata et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	An Agrypninae click beetle. The type species is C. burmitinus.
Cretophengodes^[163]	Fam. nov., Gen. et sp. nov	Valid	Li, Kundrata, Tihelka & Cai in Li et al.	Cretaceous (Albian or Cenomanian)	Burmese amber	Myanmar	A cretophengodid elateroid beetle. The type species is C. azari.
Dyscharachthis frickae^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe (Baltic Sea region)	A Dyscharachthini Eucneminae false click beetle.
Electrothroscus^[158]	Gen. et sp. nov	Valid	Li & Cai	Cretaceous Cenomanian	Burmese amber	Myanmar	A throscid false click beetle. The type species is E. yanpingae.
Entomophthalmus antennalis^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe (Baltic Sea region)	A Dirhagini Melasinae false click beetle.
Erdaia bispinulosa^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Orodotini? Macraulacinae false click beetle.
Erdaia longicornis^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Orodotini? Macraulacinae false click beetle.
Erdaia longitarsis^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Orodotini? Macraulacinae false click beetle.
Euryptychoides^[153]	Gen. et sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Euryptychini Macraulacinae false click beetle. The type species is E. magnus.
Euryptychus decoratus^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Euryptychini Macraulacinae false click beetle.
Euryptychus excavatus^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Euryptychini Macraulacinae false click beetle.
Euryptychus helmwigae^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Euryptychini Macraulacinae false click beetle.
Euryptychus sieglindae^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Euryptychini Macraulacinae false click beetle.
Euryptychus waltrautae^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An Euryptychini Macraulacinae false click beetle.
Hukawngichthyurus maha^[161]	Sp. nov	Valid	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A soldier beetle.
Isorhipis balticus^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	An Isorhipis species melasine false click beetle
Lycocerus crassepunctatus^[164]	Sp. nov	Valid	Kazantsev	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A soldier beetle.
Malthodes aliger^[164]	Sp. nov	Valid	Kazantsev	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A soldier beetle
Malthodes headsi^[165]	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber	Poland	A soldier beetle.
Malthodes jaredi^[165]	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber	Poland	A soldier beetle, a species of Malthodes.
Malthodes vladiceniensis^[166]	Sp. nov	Valid	Pintilioaie, Fanti & Ionesi	Middle Miocene	Bârnova-Muntele Formation	Romania	A soldier beetle.
Microrhagus (Emyirhagus) vetai^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Microrhagusspecies melasine false click beetle
Murcybolus^[167]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	A Burmolycini dexorine net-winged beetle. The type species is M. longiantennus.
Nematodes schwertleitae^[153]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	A Nematodini Macraulacinae false click beetle.
Neocharoides^[153]	Gen. et sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	A Nematodini? Macraulacinae false click beetle. The type species is N. balticus.
Palaeocantharis^[161]	Gen. et sp. nov	Disputed	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A soldier beetle. The type species is P. panna. Fanti & Müller (2022) considered the genus Palaeocantharis to a be a junior synonym of the genus Elektrokleinia, though the authors maintained P. panna as a distinct species within the latter genus.^[157]
Podistra guthriei^[165]	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber	Poland	A soldier beetle.
Poecilochrus balticus^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Eucnemini Eucneminae false click beetle.
Potergites brittoni^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Dendrocharini Eucneminae false click beetle.
Pseudomataeopsephus^[168]	Gen. et sp. nov	In press	Li, Huang & Cai	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A water-penny beetle. The type species is P. burmensis.
Pseudopactopus^[158]	Gen. et sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous ( Cenomanian)	Burmese amber	Myanmar	A throscid false click beetle. The type species is P. robustus.
Ptilodactyla eocenica^[169]	Sp. nov	Valid	Kundrata, Bukejs & Blank in Kundrata et al.	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	APtilodactyla species toe winged beetle.
Rhacopus balticus^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Rhacopus species eucnemid false click beetle
Rhagomicrus balticus^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Dirhagini Melasinae false click beetle.
Rheanischia^[170]	Gen. et sp. nov		Li, Chang & Muona	Early Cretaceous (Hauterivian to Aptian)	Yixian Formation	China	An Anischiinae false click beetle. The type species is R. brevicornis.
Silis curleri^[171]	Sp. nov	Valid	Fanti & Pankowski	Burdigalian	Dominican amber	Dominican Republic	A species of Silis.
Silis hegnai^[171]	Sp. nov	Valid	Fanti & Pankowski	Burdigalian	Dominican amber	Dominican Republic	A species of Silis.
Xylophilus balticus^[153]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Xylophilus species eucnemid false click beetle.

Elateriformia research

A study on the anatomy and phylogenetic relationships of Archaeolus funestus is published by Li et al. (2021).^[172]
A study on the morphological diversity of larvae of extant and fossil members of the family Brachypsectridae is published by Haug et al. (2021).^[173]

Scarabaeiformia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Goriresina^[174]	Gen. et sp. nov	Valid	Matalin, Perkovsky & Vasilenko	Late Eocene Priabonian	Rovno amber	Ukraine	A tiger beetle. The type species is G. fungifora.
Onthophagus pilauco^[175]	Sp. nov	Valid	Tello et al.	Late Pleistocene	Pilauco Bajo	Chile	A scarab beetle.

Staphyliniformia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acritus sutirca^[176]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Acritus.
Bacanius gorskii^[177]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber	Poland	A species of Bacanius.
Burmagluta^[178]	Gen. et sp. nov	Valid	Yin & Cai	Cretaceous	Burmese amber	Myanmar	A Brachyglutini Pselaphinae rove beetle. The type species is B. rougemonti.
Carinumerus^[179]	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber	Myanmar	An Onthophilinae clown beetle. The type species is C. maddisoni.
Charhyphus serratus^[180]	Sp. nov	In press	Yamamoto & Shavrin	Eocene	Baltic amber	Europe	A Phloeocharinae rove beetle.
Cretosaja^[181]	Gen. et sp. nov	In press	Sohn & Nam	Early Cretaceous	Jinju Formation	South Korea	A carrion beetle. The type species is C. jinjuensis.
Druantia^[179]	Gen. et sp. nov	Valid	Caterino	Cenomianian	Burmese amber	Myanmar	A Dendrophilinae clown beetle. The type species is D. aeterna.
Euconnus nathani^[182]	Sp. nov	Valid	Jałoszyński & Perkovsky	Eocene	Rovno amber	Ukraine	A Euconnus species Scydmaeninae rove beetle.
Eusphalerum bukejsi^[183]	Sp. nov	Valid	Shavrin in Shavrin & Kairišs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Eusphalerum species Omaliinae rove beetle.
Megabythinus^[184]	Gen. et sp. nov	Valid	Yin, Zhao & Cai	Cretaceous Cenomanian	Burmese amber	Myanmar	A Bythinini Pselaphinae rove beetle. The type species is M. spinitibialis.
Paroxyporus^[185]	Gen. et sp. nov	Valid	Cai	Early Cretaceous	Yixian Formation	China	A rove beetle. The type species is P. fossilis.
Phasmister^[179]	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber	Myanmar	An Onthophilinae clown beetle. The type species is P. cristatus.
Ponohydrochus^[186]	Gen. et sp. nov	Valid	Prokin & Strelnikova	Early Cretaceous (Hauterivian)		Russia	A water scavenger beetle. The type species is P. buryaticus.
Proptomaphaginus alleni^[187]	Sp. nov	Valid	Perreau & Haelewaters in Perreau, Haelewaters & Tafforeau	Miocene Burdigalian	Dominican amber	Dominican Republic	A Ptomaphagini round fungus beetle.
Trichophya minor^[188]	Sp. nov	Valid	Yamamoto & Newton	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A Trichophya species Trichophyinae rove beetle.
Yethiha^[179]	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber	Myanmar	A Dendrophilinae clown beetle. The type species is Y. peregrina.

other Coleopteran research

A study on the evolution of beetles from the Early Permian to Middle Triassic, evaluating the impact of the Permian–Triassic extinction event on beetle diversity, is published by Zhao et al. (2021).^[189]
Revision of Notocupes undatabdominus and N. ? multituberatus from the Lower Cretaceous of South China is published by Li, Huang & Cai (2021), who argue that both species do not belong to the genus Notocupes, transfer N. ? multituberatus to the genus Lasiosyne, and consider "N." undatabdominus to be likely a member of the family Artematopodidae.^[190]
The oldest solid wood-borer larvae reported to date (belonging to the families Buprestidae and possibly Cerambycidae) are described from the Cretaceous amber from Myanmar by Haug et al. (2021).^[191]
Mitochondrial genomes and nuclear ribosomal DNA of beetles, in some cases identifiable to species, are recovered from pack rat middens from California (United States) and Baja California (Mexico) dating up to ~ 34,355 years before present by Smith et al. (2021).^[192]

Clade Dictyoptera

New Dictyopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Afrophaga^[193]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A cockroach belonging to the family Corydiidae. Genus includes new species A. extincta.
Alienopterix mlynskyi^[194]	Sp. nov	In press	Sendi in Vršanský et al.	Cretaceous	Burmese amber	Myanmar	An alienopterid.
Alienopterix smidovae^[194]	Sp. nov	In press	Hinkelman in Vršanský et al.	Cretaceous	Burmese amber	Myanmar	An alienopterid.
Arceotermes^[195]	Fam. et gen. et sp. nov	Valid	Engel & Jiang in Jiang et al.	Cretaceous	Burmese amber	Myanmar	A termite belonging to the new family Arceotermitidae. Genus includes new species A. hospitis.
Caligoptera^[196]	Gen. et sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Ectobiidae. Genus includes new species C. hinkelmani.
Chuanblatta^[197]	Gen. et 2 sp. nov	In press	Liang et al.	Middle Jurassic	Jiulongshan Formation	China	A cockroach belonging to the family Raphidiomimidae. Genus includes new species C. huntianlingensis and C. lata.
Cuniculoblatta^[198]	Gen. et sp. nov	In press	Hinkelman	Cretaceous	Burmese amber	Myanmar	A cockroach. Genus includes new species C. brevialata .
Ectoovia^[196]	Gen. et sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Ectobiidae. Genus includes new species E. protecta.
? Elisama algeriaensis^[193]	Sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Blattulidae.
Eminespina^[199]	Gen. et sp. nov	Valid	Chen, Zhang Shi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An alienopterid. Genus includes new species E. burma.
Fractalia^[200]	Fam. et gen. et 2 sp. nov	Valid	Hinkelman & Vršanský in Vršanský et al.	Late Jurassic–Late Cretaceous (Kimmeridgian–Cenomanian)	Burmese amber	Germany Kazakhstan Myanmar	A member of Corydioidea, the type genus of the new family Fractaliidae Vršanský & Hinkelman. The type species is F. articulata Hinkelman; genus also includes new species F. aristovi Vršanský.
Fragosublatta^[201]	Gen. et sp. nov	Valid	Chen, Shih & Ren in Chen et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Corydiidae. The type species is F. pectinata.
Glyptotermes abyssinicus^[202]	Sp. nov	Valid	Bouju, Jouault & Perrichot	Early Miocene	Ethiopian amber	Ethiopia	A termite belonging to the family Kalotermitidae.
Kinneyblatta^[203]	Gen. et sp. nov	Valid	Schneider et al.	Carboniferous (Kasimovian)	Atrasado Formation	United States ( New Mexico)	A member of Blattodea belonging to the family Spiloblattinidae. The type species is K. huberi.
Kriedoblatta^[193]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Liberiblattinidae. Genus includes new species K. gondwanensis.
? Kurablattina samsonovi^[193]	Sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Liberiblattinidae.
Labradormantis^[204]	Gen. et sp. nov	Valid	Demers-Potvin et al.	Late Cretaceous (Cenomanian)	Redmond Formation	Canada	A praying mantis belonging to the family Baissomantidae. Genus includes new species L. guilbaulti.
Latiblattella basaltica^[205]	Sp. nov	In press	Vršanský, Poschmann & Vidlička	Late Oligocene		Germany	A species of Latiblattella.
Latiblattella karlgruberi^[196]	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar
Laticephalana^[206]	Gen. et sp. nov	Valid	Luo et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Umenocoleidae. Genus includes new species L. liuyani.
Lukotermes^[207]	Gen. et sp. nov	Valid	Perkovsky & Nel	Eocene (Priabonian)	Rovno amber	Ukraine	A termite belonging to the family Rhinotermitidae. Genus includes new species L. milescaput.
Meiatermes cretacicus^[208]	Sp. nov	In press	Bezerra et al.	Early Cretaceous	Crato Formation	Brazil	A termite.
Meloblatta^[193]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Mesoblattinidae. Genus includes new species M. brezinica.
Milesitermes^[209]	Gen. et sp. nov	Valid	Jouault et al.	Cretaceous	Burmese amber	Myanmar	A termite belonging to the family Mastotermitidae. Genus includes new species M. engeli.
Miniblattina^[210]	Gen. et sp. nov	In press	Sendi	Early Cretaceous (Barremian)	Lebanese amber	Lebanon	A cockroach belonging to the family Liberiblattinidae. Genus includes new species M. libera.
Mongolblatta sendii^[211]	Sp. nov	In press	Hinkelman	Cretaceous	Burmese amber	Myanmar	A member of the family Mesoblattinidae.
Morphna cenozoica^[212]	Sp. nov	In press	Šmídová, Vidlička & Wedmann	Eocene	Messel pit	Germany	A cockroach belonging to the family Blaberidae.
Morphna cretacica^[212]	Sp. nov	In press	Šmídová, Vidlička & Wedmann	Cretaceous	Burmese amber	Myanmar	A cockroach belonging to the family Blaberidae.
Nadveruzenie^[194]	Gen. et sp. nov	In press	Vršanský, Hinkelman & Sendi in Vršanský et al.	Cretaceous	Burmese amber	Myanmar	An alienopterid. Genus includes new species N. postava.
Nodosigalea simplivena^[213]	Sp. nov	In press	Šmídová	Cretaceous	Burmese amber	Myanmar	A cockroach belonging to the family Corydiidae.
Ocelloblattula margarita^[200]	Sp. nov	Valid	Koubová & Vršanský in Vršanský et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Corydioidea.
Olenablatta^[213]	Gen. et sp. nov	In press	Šmídová	Cretaceous	Burmese amber	Myanmar	A cockroach belonging to the family Olidae. Genus includes new species O. vrsanskyi.
Otazka^[193]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Mesoblattinidae. Genus includes new species O. systematicka.
Pabuonqed magna^[214]	Sp. nov	In press	Song et al.	Cretaceous	Burmese amber	Myanmar	A member of Mastotermitoidea belonging to the family Pabuonqedidae.
Paekthoblatta^[215]	Gen. et sp. nov	Valid	So et al.	Early Cretaceous	Sinuiju Formation	North Korea	A member of the family Raphidiomimidae. Genus includes new species P. coreanica.
Periplaneta (?) perialla^[216]	Sp. nov	Valid	Anisyutkin & Perkovsky	Eocene	Rovno amber	Ukraine	A cockroach, possibly a species of Periplaneta.
Petropterix fukuiensis^[217]	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation	Japan	A member of the family Umenocoleidae.
Praeblattella arcuata^[217]	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation	Japan	A member of the family Mesoblattinidae.
Praeblattella inexpecta^[217]	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation	Japan	A member of the family Mesoblattinidae.
Pseudomantina^[200]	Gen. et 2 sp. nov	Valid	Sendi & Vršanský in Vršanský et al.	Late Jurassic–Early Cretaceous (Kimmeridgian–Barremian)	Lebanese amber	Kazakhstan Lebanon	A member of the family Liberiblattinidae. The type species is P. occisor Sendi; genus also includes new species P. nigroalba Vršanský.
Reticulitermes grimaldii^[209]	Sp. nov	Valid	Jouault et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A termite, a species of Reticulitermes.
Sajda^[193]	Gen. et sp. nov	Junior homonym	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of Holocompsinae. Genus includes new species S. equatorialis. The generic name is preoccupied by Sajda Dworakowska (1981); Jiang, Xing & Li (2023) coined a replacement name Vrsanskysajda.^[218]
Spinaeblattina baekthoensis^[219]	Sp. nov	Valid	So et al.	Early Cretaceous	Sinuiju Formation	North Korea	A member of the family Mesoblattinidae.
Stavba delicata^[210]	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Liberiblattinidae.
Stavba magnoculara^[210]	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Liberiblattinidae.
Tanytermitalis^[195]	Gen. et sp. nov	Valid	Engel & Cai in Jiang et al.	Cretaceous	Burmese amber	Myanmar	A termite belonging to the family Tanytermitidae. Genus includes new species T. philetaerus.

Dictyopteran research

A study on the phylogenetic placement of the family Umenocoleidae, based on data from a new specimen of Umenocoleus sinuatus from the Lower Cretaceous Zhonggou Formation (China), is published by Liuo et al. (2021).^[220]
Description of new alienopterid nymphs from the Cretaceous Burmese amber, and a study on life history and affinities of alienopterids, is published by Luo et al. (2021).^[221]
A study on the sensory organs of Huablattula hui, and on their implications for the knowledge of the ecology of this cockroach, is published by Taniguchi et al. (2021).^[222]

Clade †Glosselytrodea

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Moltenojurina^[223]	Gen. et sp. nov	Valid	Béthoux & Anderson	Late Triassic (Carnian)	Molteno Formation	South Africa	A polycytellid Glosselytrodean. The type species is M. parva.
Polycytella rasnitsyni^[223]	Sp. nov	Valid	Béthoux & Anderson	Late Triassic (Carnian)	Molteno Formation	South Africa	A polycytellid Glosselytrodea.

Hymenopterans

New Hymenopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aethemelikertes^[224]	Gen. et sp. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A bee belonging to the tribe Melikertini. The type species is A. emunctorii.
Alivespa gracilenta^[225]	Sp. nov	In press	Wu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Vespidae.
Allommation^[226]	Fam. nov., Gen. et 3 sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family Allommationidae. Genus includes new species A. procerum, A. elongatum and A. grande.
Amelikertotes^[224]	Gen. et comb. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber Bitterfeld amber	Germany Russia ( Kaliningrad Oblast)	A bee belonging to the tribe Melikertini. The type species is "Melikertes" clypeatus Engel (2001).
Ammosphecium^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Crabroninae. Genus includes new species A. diabolicum.
Anochetus miosumatrensis^[227]	Sp. nov	Valid	Ngô-Muller, Garrouste & Nel in Ngô-Muller et al.	Miocene		Indonesia	An ant, a species of Anochetus.
Archaeovespa incompleta^[225]	Sp. nov	In press	Wu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Vespidae.
Archeonesia^[228]	Gen. et sp. nov	Valid	Tribull, Pankowski & Colombo	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Pristocerinae Bethylid. The type species is A. eocena.	Archeonesia eocena
Arkadiserphus^[229]	Gen. et sp. nov	Valid	Rasnitsyn	Late Jurassic	Karabastau Formation	Kazakhstan	A member of Proctotrupoidea belonging to the family Peleserphidae. The type species is A. leleji.
Ascogaster (Syntaphus) latitibialis^[230]	Sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Cheloninae.
Aulacicosmus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Heterogynaidae. Genus includes new species A. setosus.
Austroplebeia fujianica^[231]	Sp. nov	Valid	Engel in Engel et al.	Miocene (Langhian)	Zhangpu amber	China	A stingless bee, a species of Austroplebeia.
Burmabracon^[232]	Gen. et 2 sp. nov	Valid	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Braconidae belonging to the subfamily Protorhyssalinae. Genus includes new species B. gracilens and B. grossus.
Burmapyris^[233]	Gen. et sp. nov	Valid	Jouault, Perrichot & Nel	Cretaceous (Albian-Cenomanian)	Burmese amber	Myanmar	A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species B. azevedoi.
Burmoxyela^[234]	Gen. et sp. nov	Valid	Zheng et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A sawfly belonging to the family Syspastoxyelidae. Genus includes new species B. lii.
Burmusculus angustus^[235]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of Apocrita belonging to the family Burmusculidae.
Buserphites^[236]	Gen. et 2 sp. nov	In press	Herbert & McKellar	Cretaceous	Burmese amber	Myanmar	A member of the family Serphitidae. Genus includes new species B. applanatus and B. myanmarensis.
Chainochora^[237]	Gen. et sp. nov	Valid	Chen, van Achterberg & Hong	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is C. syntoma.
Chlorepyris deploegi^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
Chlorepyris engeli^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
Chlorepyris hopei^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
Chlorepyris jouaulti^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
Chlorepyris mckellari^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
Chlorepyris meunieri^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
Chlorepyris neli^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber Rovno amber	Russia Ukraine	A member of the family Bethylidae.
Chlorepyris perrichoti^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
Christophus^[239]	Gen. et 2 sp. nov	In press	Brazidec et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Diapriidae. Genus includes new species C. myanmarensis and C. ohmkuhnlei.
Cretastatus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Astatinae. Genus includes new species C. longevus.
Cretorussus^[240]	Gen. et sp. nov	In press	Jouault, Perrichot & Nel	Cretaceous	Burmese amber	Myanmar	A member of Orussoidea. Genus includes new species C. vilhelmseni.
Curiosivespa sennlaubi^[241]	Sp. nov	Valid	Jouault, Rasnitsyn & Nel in Jouault et al.	Early Cretaceous	Crato Formation	Brazil	A member of the family Vespidae.
Diameneura^[242]	Gen. et sp. nov	In press	Santer & Álvarez-Parra in Santer et al.	Early Cretaceous (Albian)	Escucha Formation	Spain	A member of the family Spathiopterygidae. The type species is D. marveni.
Drymomyrmex rasnitsyni^[243]	Sp. nov	Valid	Radchenko	Late Eocene	Rovno amber	Ukraine	An ant belonging to the subfamily Formicinae.
Elasmosomites arkadyleleji^[244]	Sp. nov	Valid	Belokobylskij in Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Euphorinae.
Electrofoenops exaltatus^[245]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Aulacidae.
Embolemus brachypterus^[246]	Sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Embolemidae.
Embolemus micropterus^[246]	Sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Embolemidae.
Eopelecinus marechali^[247]	Sp. nov	Valid	Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Pelecinidae.
Eoteredon^[248]	Gen. et sp. nov	Valid	Archibald, Aase & Nel	Early Eocene	Green River Formation	United States ( Wyoming)	A horntail. Genus includes new species E. lacoi.
Ephedrus rasnitsyni^[249]	Sp. nov	Valid	Davidian & Kaliuzhna in Davidian, Kaliuzhna & Perkovsky	Middle Eocene	Sakhalinian amber	Russia	A member of the family Braconidae belonging to the subfamily Aphidiinae.
Exilaulacus advenus^[245]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Aulacidae.
Exilaulacus eximius^[245]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Aulacidae.
Formica ribbeckei^[250]	Sp. nov	Valid	Radchenko & Perkovsky in Radchenko, Perkovsky & Vasilenko	Late Eocene	Rovno amber	Ukraine	An ant, a species of Formica.
Glenocephalus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family Cirrosphecidae. Genus includes new species G. mandibularis.
Gloxinius^[238]	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae; a new genus for "Epyris" bifossatus Brues (1939).
Grandixyela^[251]	Gen. et sp. nov	Valid	Zheng in Zheng et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Syspastoxyelidae. The type species is G. rasnitsyni.
Haidomelikertes^[224]	Gen. et sp. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A bee belonging to the tribe Melikertini. The type species is H. uraeus.
Haptodioctes^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family Cirrosphecidae. Genus includes new species H. apiformis.
Helosericus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psenini. Genus includes new species H. verrucosus.
Heteroichneumon^[252]	Gen. et sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae. The type species is H. rasnitsyni.
Heteropimpla pulverulenta^[252]	Sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae.
Holepyris rasnitsyni^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
Holepyris terayamai^[238]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
Holopsenella antiqua^[253]	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber	Myanmar	A member of Aculeata of uncertain phylogenetic placement.
Holopsenella burmitica^[253]	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber	Myanmar	A member of Aculeata of uncertain phylogenetic placement.
Holopsenella gothica^[253]	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber	Myanmar	A member of Aculeata of uncertain phylogenetic placement.
Hybristodryinus castaneus^[254]	Sp. nov	Valid	Olmi, Guglielmino & Chen in Olmi et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Dryinidae.
Hybristodryinus viriosus^[255]	Sp. nov	Valid	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Dryinidae.
Hybristodryinus zaifui^[254]	Sp. nov	Valid	Chen, Olmi & Perkovsky in Olmi et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Dryinidae.
Janzenella theia^[256]	Sp. nov	Valid	Bremer & Talamas in Bremer, van de Kamp & Talamas	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Janzenella.
Karataus ryonsangensis^[257]	Sp. nov	In press	Won & So	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A wasp belonging to the family Ephialtitidae.
Keratodellitha^[258]	Gen. et 3 sp. nov	Valid	Jouault et al.	Cretaceous	Burmese amber	Myanmar	A member of Evanioidea belonging to the family Othniodellithidae. Genus includes new species K. anubis, K. basilisci and K. kirin.
Kleistochora^[237]	Gen. et sp. nov	Valid	Chen, van Achterberg & Hong	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is K. dolichura.
Lelejus^[259]	Gen. et sp. nov	Valid	Radchenko & Proshchalykin	Late Eocene	Rovno amber	Ukraine	An ant belonging to the subfamily Myrmicinae. The type species is L. venustus.
Lonchodryinus groehni^[260]	Sp. nov	Valid	Olmi et al.	Late Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Dryinidae.
Mael^[238]	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae belonging to the subfamily Scleroderminae; a new genus for "Calyoza" longiceps Brues (1923).
Megalyrhyssalus^[261]	Gen. et sp. nov	Valid	Belokobylskij & Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae. Genus includes new species M. clavicornis.
Megarolium^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Bembicinae. Genus includes new species M. paradoxum.
Merascylla^[238]	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Bethylidae belonging to the subfamily Pristocerinae; a new genus for "Epyris" atavellus Cockerell (1920).
Mesoserphites^[236]	Gen. et 5 sp. nov	In press	Herbert & McKellar	Cretaceous	Burmese amber	Myanmar	A member of the family Serphitidae. Genus includes new species M. annulus, M. giganteus, M. engeli, M. scutatus and M. viraneacapitis.
Meteorus arasnitsyni^[230]	Sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Meteorus.
Microtypus eocenus^[230]	Sp. nov	Valid	Belokobylskij et al.	Eocene	Rovno amber	Ukraine	A member of the family Braconidae belonging to the subfamily Microtypinae.
Mintara^[239]	Gen. et sp. nov	In press	Brazidec et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Diapriidae. Genus includes new species M. parva.
Odontosericus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psenini. Genus includes new species O. burmensis.
Orthosyntexis^[262]	Gen. et 2 sp. nov	Valid	Gao, Engel, Shih, Gao in Gao et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Anaxyelidae. The type species is O. elegans; genus also includes O. thanti.
Palaeocolastes^[230]	Gen. et sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Exothecinae. The type species is P. bruesi.
Paleoaulacus^[263]	Gen. et sp. nov	Valid	Jouault & Nel	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Aulacidae. Genus includes new species P. minutus.
Paleoxyela^[264]	Gen. et sp. nov	Valid	Jouault, Aase & Nel	Eocene (Chadronian)	Florissant Formation	United States ( Colorado)	A member of the family Xyelidae. The type species is P. nearctica.
Peleserphus qufuensis^[265]	Sp. nov	In press	Yu et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Peleserphidae.
Pheidole pauchil^[266]	Sp. nov	Valid	Varela-Hernández & Riquelme	Miocene	Mexican amber	Mexico	An ant, a species of Pheidole.
Pinguixyela^[234]	Gen. et comb. nov	Valid	Zheng et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A sawfly belonging to the family Syspastoxyelidae; a new genus for "Syspastoxyela" pinguis.
Platyxyela tenuis^[267]	Sp. nov	Valid	Zheng et al.	Middle Jurassic (Callovian-Bathonian boundary)	Jiulongshan Formation	China	A member of the family Xyelidae.
Pompilopterus ryonsangensis^[257]	Sp. nov	In press	Won & So	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A wasp belonging to the family Angarosphecidae.
Pristinopterus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Bembicinae. Genus includes new species P. spectabilis.
Pristopsen^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psenini. Genus includes new species P. obscurus.
Prosyntexis sennlaubi^[268]	Sp. nov	Valid	Jouault & Nel	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Sepulcidae.
Protopyris^[233]	Gen. et sp. nov	Valid	Jouault & Nel in Jouault, Perrichot & Nel	Early Cretaceous (Albian)	Hkamti amber	Myanmar	A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species P. myanmarensis.
Prototeleia^[269]	Gen. et sp. nov	Valid	Talamas et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Platygastridae. The type species is P. kleio.
Proxyelia^[264]	Gen. et sp. nov	Valid	Jouault, Aase & Nel	Eocene (Ypresian)	Green River Formation	United States ( Wyoming)	A member of the family Xyelidae. The type species is P. pankowskii.
Ptilocosmus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Heterogynaidae. Genus includes new species P. corniculatus.
Ramageoptera^[270]	Gen. et sp. nov	Valid	Jouault & Brazidec	Cretaceous	Burmese amber	Myanmar	A member of the family Bethylidae belonging to the subfamily Protopristocerinae. Genus includes new species R. platycephala.
Rasnichneumon^[252]	Gen. et 2 sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae. The type species is R. alexandri; genus also includes R. gracilis.
Rasnitsynum^[271]	Gen. et sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Dryinidae. Genus includes new species R. burmense.
Rogichneumon^[252]	Gen. et sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae. The type species is R. braconidicus.
Rovenosa alexrasnitsyni^[272]	Sp. nov	Valid	Manukyan in Manukyan & Zhindarev	Late Eocene	Prussian Formation (Baltic amber)	Russia ( Kaliningrad Oblast)	A member of the family Ichneumonidae.
Rovenosa khalaimi^[272]	Sp. nov	Valid	Manukyan in Manukyan & Zhindarev	Late Eocene	Prussian Formation (Baltic amber)	Russia ( Kaliningrad Oblast)	A member of the family Ichneumonidae.
Sakhalinencyrtus^[273]	Gen. et sp. nov	Valid	Simutnik in Simutnik, Perkovsky & Vasilenko	Middle Eocene	Sakhalinian amber	Russia	A member of the family Encyrtidae. The type species is S. leleji.
Sakhalinoctonus^[274]	Gen. et sp. nov	Valid	Davidian in Davidian, Manukyan & Belokobylskij	Middle Eocene	Sakhalinian amber	Russia	A member of the family Braconidae belonging to the subfamily Aphidiinae. Genus includes new species S. alexrasnitsyni.
Scleroxyela^[267]	Gen. et sp. nov	Valid	Zheng et al.	Middle Jurassic (Callovian-Bathonian boundary)	Jiulongshan Formation	China	A member of the family Xyelidae. The type species is S. daohugouensis.
Spalangiopelta darlingi^[275]	Sp. nov	Valid	Moser in Moser et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Pteromalidae.
Spalangiopelta semialba^[275]	Sp. nov	Valid	Moser in Moser et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Pteromalidae.
Spheciellus^[226]	Fam. nov., Gen. et sp. nov.	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family Spheciellidae. Genus includes new species S. aenigmaticus.
Stephanorhyssalus^[261]	Gen. et sp. nov	Valid	Belokobylskij & Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae. Genus includes new species S. longiscapus.
Tetragonula florilega^[231]	Sp. nov	Valid	Engel in Engel et al.	Miocene (Langhian)	Zhangpu amber	China	A stingless bee, a species of Tetragonula.
Thigmocosmus^[226]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Heterogynaidae. Genus includes new species T. longicornis.
Tyrannomecia^[276]	Gen. et sp. nov	In press	Jouault & Nel	Paleocene	Menat Formation	France	An ant belonging to the stem group of the subfamily Myrmeciinae. Genus includes new species T. inopinata.
Vespula? hassiaca^[277]	Sp. nov	In press	Abels & Wedmann	Eocene	Messel pit	Germany	A member of the family Vespidae.
Xeris dorbnikensis^[278]	Sp. nov	Valid	Manukyan in Manukyan & Smirnova	Eocene	Baltic amber	Europe (Baltic Sea region)	A horntail. Originally described as a species of Xeris; Archibald & Rasnitsyn (2022) transferred it to the genus Urocerus.^[279]
Zigrasimecia goldingot^[280]	Sp. nov	In press	Zhuang et al.	Cretaceous	Burmese amber	Myanmar	An ant in the tribe Sphecomyrmini.

Hymenopteran Research

Guo et al. (2021) report the discovery of two queen ant specimens from Burmese amber, belonging to the species Haidomyrmex cerberus and including the first known alate queen in Haidomyrmex.^[281]
Cooccurrence of ants belonging to extant Holarctic genus Lasius and a tropical weaver ant in the same piece of amber is reported from the Bitterfeld amber (Germany) by Radchenko & Perkovsky (2021).^[282]
Original claims that both Melittosphex burmensis and Discoscapa apicula were primitive bees have been rejected by Rosa & Melo, who instead consider the former to be an aculeate wasp of uncertain affinity, and the latter to belong within the crabronid wasp subfamily Crabroninae, a lineage not directly related to bees.^[226]

Clade Neuropterida

Neuropterans

New Neuropteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Ankyloleon^[283]	Gen. et sp. nov	Valid	Badano, Haug & Cerretti in Badano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An early representative of the clade Myrmeleontiformia. The type species is A. caudatus.
Astioberotha coutreti^[284]	Sp. nov	In press	Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Rhachiberothidae belonging to the subfamily Paraberothinae.
Burmotachinymphes^[285]	Gen. et sp. nov	Valid	Cao, Wang & Liu	Cretaceous	Burmese amber	Myanmar	A member of the family Mesochrysopidae. Genus includes new species M. bilobata.
Calobabinskaia^[286]	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Babinskaiidae. The type species is C. xiai.
Fuscopolystoechotes^[287]	Gen. et sp. nov	Valid	Xu, Zheng, Shih & Wang in Xu et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Ithonidae. Genus includes new species F. reticulatus.
Karenina cuneiformis^[288]	Sp. nov	Valid	Nakamine	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Mesochrysopidae.
Lichenipolystoechotes fenestratus^[287]	Sp. nov	Valid	Xu, Fang, Shih & Wang in Xu et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Ithonidae.
Osmylidia donnae^[289]	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Quilchena site	Canada British Columbia	A protosmyline Osmylid lance lacewing
Osmylidia glastrai^[289]	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Klondike Mountain Formation	United States Washington	A protosmyline Osmylid lance lacewing
Osmylidia picta^[289]	Comb. nov	Valid	(Hagen)	Priabonian	Baltic amber	Europe	A protosmyline Osmylid lance lacewing Moved from "Protosmylus" picta (1913)
Osmylidia taliae^[289]	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Green River Formation Parachute Creek Member	United States Colorado	A protosmyline Osmylid lance lacewing
Osmyloberotha^[290]	Gen. et sp. nov	Valid	Khramov	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A beaded lacewing. Genus includes new species O. simpla.
Paradoxoberotha^[291]	Gen. et sp. nov	In press	Nakamine et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Rhachiberothidae. Genus includes new species P. chimaera.
Paraneliana^[292]	Gen. et sp. nov	Valid	Jouault & Nel	Early Cretaceous	Crato Formation	Brazil	A member of the family Babinskaiidae. Genus includes new species P. sennlaubi.
Rasnitsyneura^[293]	Gen. et comb. nov	Valid	Makarkin, Ansorge & Khramov	Early Jurassic (Toarcian)		Germany	A member of the family Ithonidae sensu lato belonging to the subfamily Epigambriinae. The type species is "Prohemerobius" aequabilis Bode (1953).
Sinonothochrysa^[294]	Gen. et sp. nov	In press	Huang & Nel in Huang et al.	Paleocene	Buxin Formation	China	A member of the family Chrysopidae. Genus includes new species S. zhangi
Stenobabinskaia^[286]	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Babinskaiidae. The type species is S. punctata.
Xiaobabinskaia^[286]	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Babinskaiidae. The type species is X. lepidotricha.

Neuropteran research

Review of the known record of spoon-winged lacewing larvae is published Haug, Haug & Haug (2021), who report two specimens from the Cretaceous amber from Myanmar which might represent the first known larvae of members of the subfamily Nemopterinae in the fossil record.^[295]
New lacewing larvae with elongated mouthparts are described from the Cretaceous amber from Myanmar by Haug et al. (2021), who also review the morphological diversity of fossil lacewing larvae.^[296]

Raphidiopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Archiinocellia protomaculata^[297]	comb nov	valid	(Engel)	Lutetian	Green River Formation	USA Colorado	A raphidiid snakefly. Moved from Agulla protomaculata (2011)^[298]	Archiinocellia protomaculata
Electrobaissoptera liui^[299]	Sp. nov	In press	Jouault, Engel & Nel	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A baissopterid snakefly.
Megaraphidia antiquissima^[297]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Tranquille Formation	Canada British Columbia	A raphidiid snakefly.
Megaraphidia hopkinsi^[297]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Allenby Formation	Canada British Columbia	A raphidiid snakefly.
Megaraphidia klondika^[297]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Klondike Mountain Formation	United States Washington	A raphidiid snakefly.
Megaraphidia ootsa^[297]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	"Unnamed unit" Ootsa Group	Canada British Columbia	A raphidiid snakefly.
Mesoraphidia baekthoensis^[300]	Sp. nov	In press	Ri et al.	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A member of the family Mesoraphidiidae.
Mesoraphidia ryonsangensis^[300]	Sp. nov	In press	Ri et al.	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A member of the family Mesoraphidiidae.

Clade †Palaeodictyopteroidea

†Megasecoptera

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Anhuihymen^[301]	Gen. et sp. nov	In press	Huang & Nel in Huang et al.	Middle Permian	Yinping Formation	China	A protohymenid Megasecoptera. The type species is A. medianelongata.

†Palaeodictyoptera

New Palaeodictyopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Dictyoneura goujonorum^[302]	Sp. nov	In press	Logghe et al.	Carboniferous-Permian (Gzhelian-Asselian)		France	A member of Palaeodictyoptera.
Sandiella herbigi^[303]	Sp. nov	In press	Brauckmann, Herd & Leipner	Carboniferous (Moscovian)		Germany	A member of Palaeodictyoptera belonging to the family Eugereonidae.

Palaeodictyopteran research

A study on the anatomy of fossil specimens described as consecutive series of different larval stages of palaeodictyopteran species Tchirkovaea guttata and Paimbia fenestrata, evaluating their implications for the knowledge of wing development in palaeodictyopterans, is published by Rosová, Sinitshenkova & Prokop (2021).^[304]

Clade Palaeoptera

Ephemeroptera

New Ephemeropteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Astraeoptera^[305]	Gen. et sp. nov	Valid	Brandão et al.	Early Cretaceous	Crato Formation	Brazil	A mayfly of uncertain phylogenetic placement. Genus includes new species A. cretacica.
Burmella^[306]	Gen. et 2 sp. nov	Valid	Godunko, Martynov & Staniczek	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A vietnamellid mayfly. The type species is B. paucivenosa; genus also includes B. clypeata.
Protoligoneuria heloisae^[307]	Sp. nov	In press	Storari et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil	A hexagenitid mayfly.
Triassodotes rasnitsyni^[308]	Sp. nov	Valid	Sinitshenkova	Triassic		Russia	A misthodotid mayfly.

Ephemeropteran research

Sroka et al. (2021) provide new information on the morphology of Permian stem-mayflies Misthodotes sharovi and M. zalesskyi from the Tshekarda locality (Kungurian Koshelevka Formation; Perm Krai, Russia), and infer the life history traits of both the adult and larval stages of these insects.^[309]
Possible underwater mayfly burrows are described from the Turonian Ferron Sandstone (Utah, United States) by King et al. (2021), who interpret this finding as evidence that burrowing mayflies adapted to domicile filter-feeding during or prior to the Turonian, and name a new ichnospecies Glossifungites gingrasi.^[310]

Odonatoptera

Odonata

New Odonatan taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Chalcolestes tibetensis^[311]	Sp. nov	In press	Xia et al.	Late Eocene	Nima Basin	China	A species of Chalcolestes.
Cretaphylolestes^[312]	Gen. et sp. nov	In press	Huang et al.	Early Cretaceous (Aptian)	Shouchang Formation	China	A damselfly belonging to the family Synlestidae. Genus includes new species C. cretacicus.
Dakotaeschnidium^[313]	Gen. et sp. nov	Valid	Nel	Late Cretaceous (Maastrichtian)	Fox Hills Formation	United States ( South Dakota)	A dragonfly belonging to the family Aeschnidiidae. Genus includes new species D. spedeni.
Dysagrion pruettae^[314]	Sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A member of the family Dysagrionidae.	Dysagrion pruettae
Dysagrionites^[314]	Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A collective group name for any fossil species referable to the Dysagrioninae whose orthotaxon generic affinity is unclear. Includes new species D. delinei.
Epilestes rasnitsyni^[315]	Sp. nov	Valid	Felker & Vasilenko	Permian		Russia	A damselfly belonging to the family Permagrionidae.
Jingguaeshna^[316]	Gen. et sp. nov	Valid	Zheng et al.	Miocene		China	A dragonfly belonging to the group Aeshnoidea, possibly a member of the family Telephlebiidae. Genus includes new species J. taoae.
Marado^[317]^[318]	Gen. et sp. nov	Valid	Petrulevičius	Paleocene	Maíz Gordo Formation	Argentina	A damselfly belonging to the family Coenagrionidae. The type species is M. marado.
Moltenophlebia^[319]	Gen. et sp. nov	Valid	Deregnaucourt et al.	Triassic	Molteno Formation	South Africa	A member of the stem group of Odonata, belonging to the group Zygophlebiida within the Triadophlebiomorpha. Genus includes new species M. lindae.
Okanagrion^[314]	Gen. et 8 sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A member of the family Dysagrionidae. The type species is O. threadgillae; genus also includes new species O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum and O. worleyae.	Okanagrion threadgillae
Okanopteryx^[314]	Gen. et 3 sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Tranquille Formation	Canada ( British Columbia)	A member of the family Dysagrionidae. The type species is O. macabeensis; genus also includes new species O. fraseri and O. jeppesenorum.
Phyonganpodagrion^[320]	Gen. et sp. nov	In press	So & Won	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A damselfly in the family Megapodagrionidae. Genus includes new species P. ryonsangae.
Protohemiphlebia^[321]	Fam. et gen. et 2 sp. nov	In press	Zheng, Jarzembowski & Nel in Zheng et al.	Cretaceous	Burmese amber	Myanmar	A stem hemiphlebioid damselfly belonging to the new family Protohemiphlebiidae. Genus includes new species P. zhangi and P. meiyingae.
Pulchrairina^[322]	Gen. et sp. nov	In press	Martynov, Vasilenko & Perkovsky	Late Eocene	Rovno amber	Ukraine	A damselfly belonging to the group Coenagrionoidea. Genus includes new species P. electra.
Republica^[323]	Gen. et sp. nov	Valid	Archibald & Cannings	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A euphaeid damselfly. Type species is R. weatbrooki. Not to be confused with the plant genus Republica also from the formation.	Republica weatbrooki
Stenodiafanus^[314]	Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A dysagrionid odonate. Type species is S. westersidei.	Stenodiafanus westersidei
Stenophlebia ryonsangensis^[324]	Sp. nov	In press	Won, So & Kim	Early Cretaceous	Sinuiju Formation	North Korea	A stenophlebiid odonate.
Whetwhetaksa^[314]	Fam. Nov., Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States	The type genus of Whetwhetaksidae, a new family of uncertain placement. The type species is W. millerae.	Whetwhetaksa millerae

Odonate research

Archibald et al. (2021) establish a new, extinct suborder of Odonata named Cephalozygoptera, close to but distinct from damselflies, composed of the families Dysagrionidae, Sieblosiidae and possibly Whetwhetaksidae;^[314] their study is subsequently criticized by Nel & Zheng (2021), who consider evidence presented by Archibald et al. to be insufficient to support a new clade, and reject the suborder Cephalozygoptera as unfounded.^[325]^[326]

†Protozygoptera

Name	Novelty	Status	Authors	Age	Country	Notes
Kennedya kedrovkensis^[327]	Sp. nov	Valid	Felker	Early Triassic	Russia	A member of Protozygoptera belonging to the family Kennedyidae.
Kennedya suchonensis^[328]	Sp. nov	Valid	Felker	Late Permian	Russia
Progoneura kemerovensis^[327]	Sp. nov	Valid	Felker	Early Triassic	Russia	A member of Protozygoptera belonging to the family Kennedyidae.

Other odonatopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Brunellopteron^[329]	Gen. et sp. nov	Valid	Béthoux, Deregnaucourt & Norrad in Béthoux et al.	Carboniferous (Moscovian)	Sunbury Creek Formation	Canada ( New Brunswick)	A member of Neodonatoptera of uncertain phylogenetic placement. The type species is B. norradi.
Osnabruggiala^[330]	Gen. et sp. nov	Valid	Zessin, Brauckmann & Gröning	Carboniferous (Pennsylvanian)	Osnabrück Formation	Germany	Probably a basal member of Odonatoptera. The type species is O. seppelti.

Clade †Paoliidea

†Paoliida

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Avionblattinopsis^[331]	Gen. et sp. nov	In press	Quispe et al.	Carboniferous (Moscovian)		France	A blattinopsid Paoliidan. The type species is A. oudardi.
Blattinopsis indefinitus^[332]	Sp. nov	Valid	Rasnitsyn & Aristov	Permian (Kungurian)		Russia ( Krasnoyarsk Krai)	A Blattinopsidae Paoliidan
Glaphyrophlebia borea^[333]	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation	Russia ( Komi Republic)	A Blattinopsid.
Glaphyrophlebia iva^[334]	Sp. nov	Valid	Aristov & Rasnitsyn	Middle Permian		Russia	A member of the family Blattinopsidae.
Glaphyrophlebia komia^[333]	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation	Russia ( Komi Republic)	A member of the family Blattinopsidae.
Glaphyrophlebia vorkutensis^[333]	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation	Russia ( Komi Republic)	A member of the family Blattinopsidae.

Clade Paraneoptera

Hemipterans

New Hemipteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acrotiara^[335]	Trib. et gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cixiidae; the type genus of the new tribe Acrotiarini. The type species is A. multigranulata Luo & Bourgoin.
Aneurus damzeni^[336]	Sp. nov	Valid	Heiss in Heiss, Kairišs & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Aneurus.
Aphelicophontes^[337]	Gen. et sp. nov	Valid	Swanson et al.	Eocene	Green River Formation	United States	A member of the family Reduviidae belonging to the subfamily Harpactorinae. Genus includes new species A. danjuddi.
Baltiomiris^[338]	Gen. et sp. nov	Valid	Kim, Chérot & Jung	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Miridae. The type species is B. herczeki.
Barremixius^[335]	Gen. et comb. nov	In press	Bourgoin & Szwedo in Luo et al.	Early Cretaceous	Weald Clay	United Kingdom	A member of the family Cixiidae; a new genus for "Cixius" petrinus Fennah.
Beaconiella cincta^[339]	Sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation	Kyrgyzstan	A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.
Beaconiella pulchra^[339]	Sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation	Kyrgyzstan	A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.
Burmacader bicoloripennis^[340]	Sp. nov	Valid	Souma, Yamamoto & Takahashi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Tingidae.
Burmacader grandis^[341]	Sp. nov	Valid	Heiss & Guilbert	Cretaceous	Burmese amber	Myanmar	A member of the family Tingidae.
Burmala^[342]	Gen. et sp. nov	Valid	Liu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Malmopsyllidae. Genus includes new species B. liaoyaoi.
Burmaprosbole^[343]	Gen. et sp. nov	Valid	Qiao et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Tettigarctidae. Genus includes new species B. nervosa.
Burmavianaida^[344]	Gen. et sp. nov	Valid	Souma, Yamamoto & Takahashi	Cretaceous (Albian or Cenomanian)	Burmese amber	Myanmar	A member of the family Tingidae. The type species is B. anomalocapitata.
Cretamystilus^[345]	Gen. et sp. nov	Valid	Kim, Lim & Jung	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Miridae belonging to the tribe Mecistoscelini. Genus includes new species C. herczeki.
Cretodorus rostellatus^[346]	Sp. nov	In press	Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Mimarachnidae.
Cretomultinervis^[347]	Gen. et sp. nov	Valid	Fu & Huang	Cretaceous	Burmese amber	Myanmar	A member of the family Sinoalidae. Genus includes new species C. burmensis.
Dachibangus hui^[348]	Sp. nov	Valid	Zhang, Yao & Pang in Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Mimarachnidae. Originally described as a species of Dachibangus, but subsequently transferred to the genus Xiaochibangus.^[349]
Delphitiara^[335]	Gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is D. tibiocoronata Luo & Bourgoin.
Dorytocus jiaxiaoae^[350]	Sp. nov	Valid	Song, Szwedo & Bourgoin in Song et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A planthopper.
Duyana^[351]	Gen. et sp. nov	Valid	Chen et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A leafhopper belonging to the subfamily Ledrinae. Genus includes new species D. electronica.
Electromyiomma herczeki^[352]	Sp. nov	Valid	Kim & Jung	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Miridae.
Eospinosus^[353]	Gen. et 2 sp. nov	Valid	Wedmann et al.	Eocene	Messel pit	Germany United States ( Colorado)	A member of the family Pentatomidae. The type species is E. peterkulkai; genus also includes E. greenriverensis.
Ferriantenna^[354]	Gen. et sp. nov	Valid	Cumming & Le Tirant	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Coreidae belonging to the subfamily Coreinae. The type species is F. excalibur.
Hallex martinsnetoi^[355]	Sp. nov	In press	Monferran et al.	Early Cretaceous	Crato Formation	Brazil	A leafhopper.
Hypsotsijilia^[356]	Gen. et sp. nov	Valid	Cifuentes-Ruiz & Brailovsky in Cifuentes-Ruiz et al.	Early Miocene	La Quinta Formation (Mexican amber)	Mexico	A member of the family Schizopteridae. The type species is H. bretoni.
Inoderbe^[357]	Fam. et gen. et sp. nov	Valid	Shcherbakov & Emeljanov	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A planthopper belonging to the new family Inoderbidae. The type species is I. rapunzel.
Juroala pulchra^[358]	Sp. nov	Valid	Fu & Huang	Late Jurassic	Daohugou Beds	China	A member of the family Sinoalidae.
Karatauella^[359]	Nom. nov	Valid	Doweld	Jurassic (Bathonian-Oxfordian)		Kazakhstan	A member of the family Protopsyllidiidae; a replacement name for Carpenterella Becker-Migdisova (1968).
Latidorsum^[360]	Gen. et sp. nov		Wang, Tang & Yao in Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Tingidae. The type species is L. carinbifarium.
Microtingis^[361]	Gen. et sp. nov	Junior synonym	Heiss & Golub	Cretaceous	Burmese amber	Myanmar	A member of the family Tingidae. Genus includes new species M. leptosoma. Subsequently Heiss & Golub (2021) considered this species to be a junior synonym of Burmavianaida anomalocapitata.^[362]
Multistria^[348]	Gen. et sp. nov	Valid	Zhang, Yao & Pang in Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Mimarachnidae. The type species is M. orthotropa.
Paleocader balticus^[363]	Sp. nov	Valid	Heiss & Golub	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Tingidae.
Paleocader pulchellus^[363]	Sp. nov	Valid	Heiss & Golub	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Tingidae.
Paleocader rovnensis^[363]	Sp. nov	Valid	Heiss & Golub	Eocene	Rovno amber	Ukraine	A member of the family Tingidae.
Parasinalda sukachevae^[364]	Sp. nov	Valid	Golub, Perkovsky & Vasilenko	Late Eocene	Rovno amber	Ukraine	A member of the family Tingidae.
Pentacarinus^[335]	Gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is P. kachinensis Luo & Bourgoin.
Pseudocaulisoculus^[365]	Gen. et sp. nov	Valid	Kóbor & Roca-Cusachs	Cretaceous	Burmese amber	Myanmar	A member of the family Yuripopovinidae. Genus includes new species P. longicornis.
Pubivetanthocoris^[366]	Gen. et sp. nov	In press	Tang, Wang & Yao in Tang et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Vetanthocoridae. Genus includes new species P. carinalis.
"Pyrrhocoris" rottensis^[367]	Nom. nov	Valid	Kment	Late Oligocene		Germany	A member of Lygaeoidea of uncertain phylogenetic placement; a replacement name for Pyrrhocoris tibialis Statz & Wagner (1950).
Rovnoxestus^[368]	Gen. et sp. nov	Valid	Dietrich, Dmitriev & Perkovsky	Late Eocene	Rovno amber	Ukraine	A leafhopper. Genus includes new species R. rasnitsyni.
Saltissus fennahi^[369]	Sp. nov	In press	Luo, Liu & Jarzembowski in Luo et al.	Early Cretaceous	Weald Clay	United Kingdom	A member of the family Mimarachnidae.
Scytinoptera tongchuanensis^[370]	Sp. nov	In press	Zhang, Zheng & Zhang in Zhang et al.	Middle Triassic	Tongchuan Formation	China	A member of the family Scytinopteridae.
Sinogranulus^[371]	Gen. et sp. nov	In press	Zhang, Chen & Zhang	Middle Triassic	Tongchuan Formation	China	A member of Scytinopteroidea belonging to the family Granulidae. Genus includes new species S. qishuiheensis.
Spinonympha^[372]	Gen. et sp. nov	Valid	Luo, Wang & Jarzembowski	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A planthopper of uncertain phylogenetic placement. The type species is S. shcherbakovi.
Szeiinia^[373]	Fam. nov., Gen. et sp. nov	Valid	Zhang et al.	Late Triassic	Yanchang Formation	China	A planthopper in the new family Szeiiniidae. Genus includes new species S. huanglongensis
Tennentsia orientalis^[374]	Sp. nov	In press	Fu, Azar & Huang	Middle Triassic	Yanchang Formation	China	A member of Cicadomorpha belonging to the family Dysmorphoptilidae.
Triknightia^[339]	Gen. et sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation	Kyrgyzstan	A member of Cicadomorpha belonging to the superfamily Scytinopteroidea and the family Paraknightiidae. The type species is T. mira.
Tuthillia maculosa^[375]	Sp. nov	Valid	Poinar & Brown		Dominican amber	Dominican Republic	A member of Psylloidea.
Valdicopis^[376]	Gen. et sp. nov	Valid	Li, Chen & Jarzembowski	Early Cretaceous		United Kingdom	A member of the family Procercopidae. Genus includes new species V. tonyi.
Youngeewa simillima^[377]	Sp. nov	Valid	Gębicki & Walczak in Gębicki, Walczak & Zmarzły	Eocene	Baltic amber	Europe (Baltic Sea region)	A leafhopper belonging to the subfamily Mileewinae.

Hemipteran research

Grimaldi & Vea (2021) reinterpret Mesophthirus engeli as a scale insect, and reject the interpretation of this insect as an ectoparasite.^[378]
Redescription of the holotype specimen of Paranoika placida is published by Moura-Júnior et al. (2021), who transfer this species to the genus Lethocerus, and consider this species to be the senior synonym of Lethocerus vetus.^[379]
A fulgorid specimen, representing the oldest occurrence of this family in the fossil record in Asia, is described from the Eocene Lunpola Basin (Qinghai-Tibetan Plateau, China) by Xu et al. (2021).^[380]
Revision of the fossil record of Fulgoromorpha and Cicadomorpha from the Jurassic and Cretaceous of eastern Asia is published by Chen (2021).^[381]

Psocodeans

New Psocodean taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Burmesopsocus^[382]	Gen. et sp. nov	Valid	Yoshizawa in Yoshizawa & Yamamoto	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A member of Homilopsocidea of uncertain phylogenetic placement. The type species is B. lienhardi.
Cormopsocus baleoi^[383]	Sp. nov	Valid	Hakim, Azar & Huang	Cretaceous	Burmese amber	Myanmar	A member of Trogiomorpha belonging to the family Cormopsocidae.
Cormopsocus neli^[384]	Sp. nov	Valid	Hakim, Azar & Huang in Hakim et al.	Cretaceous	Burmese amber	Myanmar	A member of Trogiomorpha belonging to the family Cormopsocidae.
Empherium^[385]	Gen. et sp. nov	Valid	Hakim, Huang & Azar	Late Cretaceous		Russia	A member of Trogiomorpha belonging to the family Empheriidae. Genus includes new species E. rasnitsyni.
Eolepinotus zherikhini^[385]	Sp. nov	Valid	Hakim, Huang & Azar	Late Cretaceous		Russia	A member of Trogiomorpha belonging to the family Trogiidae.
Heliadesdakruon^[386]	Gen. et sp. nov	Valid	Cumming & Le Tirant	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Trogiomorpha belonging to the family Archaeatropidae. The type species is H. morganae.
Longiglabellus^[387]	Gen. et 2 sp. nov	In press	Wang, Li & Yao in Wang et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Cormopsocidae. Genus includes new species L. edentatus and L. pedhyalinus.
Palaeosiamoglaris hammanaensis^[388]	Sp. nov	In press	Hakim, Huang & Azar	Early Cretaceous (Barremian)	Lebanese amber	Lebanon	A member of Psocodea belonging to the family Prionoglarididae.
Palaeosiamoglaris hkamtiensis^[389]	Sp. nov	In press	Jouault et al.	Early Cretaceous	Hkamti amber	Myanmar	A member of Psocodea belonging to the family Prionoglarididae.
Psyllipsocus myanmarensis^[389]	Sp. nov	In press	Jouault et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.
Psyllipsocus yangi^[390]	Sp. nov	Valid	Liang & Liu	Cretaceous	Burmese amber	Myanmar	A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.
Stimulopsocus^[391]	Gen. et sp. nov	In press	Liang & Liu	Cretaceous	Burmese amber	Myanmar	A member of the family Cormopsocidae. Genus includes new species S. jiewenae.

Psocodean research

Four bark lice carrying sand granules and organic material atop their back are described from the Cretaceous amber from Myanmar by Kiesmüller et al. (2021), who interpret this finding as the first fossil evidence of masking behaviour in Cretaceous representatives of Psocodea.^[392]

Clade Perlidea

Dermapterans

New Dermapteran

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Aglyptoderma^[393]	Gen. et 2 sp. nov	Valid	Xiong et al.	Middle Jurassic	Jiulongshan Formation	China	A semenoviolid eodermapteran earwig. Genus includes the new species A. elongatum and A. cylindratum.
Aneuroderma^[394]	Gen. et sp. nov	Valid	Xiong, Engel & Ren in Xiong et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Protodiplatyidae. The type species is A. oiodes.
Dacryoderma^[395]	Gen. et comb. nov	Valid	Engel	Early Jurassic (Sinemurian)	Charmouth Mudstone Formation	United Kingdom	A dermapterid earwig. The type species is "Brevicula" teres Tihelka (2019).
Eminepygia^[396]	Gen. et sp. nov	Valid	Chen & Zhang	Cenomanian	Burmese amber	Myanmar	A pygidicranid earwig. Genus includes new species E. myanmarensis.
Sinopalaeodermata concavum^[394]	Sp. nov	Valid	Xiong, Engel & Ren in Xiong et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Dermapteridae.

Plecoptera

New Plecopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Balticonemoura^[397]	Gen. et sp. nov	In press	Chen	Eocene	Baltic amber	Europe (Baltic Sea region)	A nemourid stonefly. Genus includes new species B. bulbosus.
Billoperla^[398]	Gen. et sp. nov	In press	Chen & Xu	Cretaceous	Burmese amber	Myanmar	A perlid stonefly belonging to the family. Genus includes new species B. starki.
Isoperla baltica^[399]	Sp. nov	Valid	Jouault et al.	Eocene	Baltic amber	Europe (Baltic Sea regon)	A perlodid stonefly.

Other insects

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Anakitala^[400]	Gen. et sp. nov	Valid	Aristov	Late Permian		Russia	A mesorthopterid Eoblattidan Polyneopteran. The type species is A. intermittendia.
Baharellinus elegans^[400]	Sp. nov	Valid	Aristov	Late Permian		Russia	A member of Polyneoptera belonging to the group Eoblattida and the family Blattogryllidae.
Belmophenopterum rasnitsyni^[401]	Sp. nov	Valid	Storozhenko & Aristov	Triassic (Ladinian-Carnian)	Madygen Formation	Kyrgyzstan	A member of the family Mesorthopteridae.
Cabarzopterum^[402]	Gen. et sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of Grylloblattida of uncertain phylogenetic placement. Genus includes new species C. magnificus.
Delopterinus alsenzensis^[403]	Sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin	Germany	A miomopteran belonging to the family Palaeomanteidae.
Delopterum novokshonovi^[403]	Sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin	Germany	A miomopteran belonging to the family Palaeomanteidae.
Elmocladus^[404]	Gen. et comb. nov	Valid	Rasnitsyn & Aristov	Early Permian		United States	A member of Polyneoptera belonging to the group Eoblattida and the family Tococladidae. The type species is "Opisthocladus" strictus Carpenter (1976).
Galloembia^[405]	Gen. et sp. nov	Valid	Falières, Engel & Nel	Earliest Eocene	Oise amber	France	A webspinner belonging to the family Embiidae. The type species is G. raholai.
Gnethoda odontophora^[406]	Sp. nov	In press	Lai et al.	Cretaceous	Burmese amber	Myanmar	A webspinner belonging to the family Clothodidae.
Kamopanorpa rasnitsyni^[407]	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin	Russia	A member of the (stem-trichopteran or stem-amphiesmenopteran) family Microptysmatidae.
Kamopanorpa rotunda^[407]	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin	Russia	A member of the family Microptysmatidae.
Kamopanorpa sivchikovi^[407]	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin	Russia	A member of the family Microptysmatidae.
Liomopterites sperbersbachensis^[402]	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of Grylloblattida belonging to the family Liomopteridae.
Liomopterum fuscatum^[402]	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of Grylloblattida belonging to the family Liomopteridae.
Madygembia^[408]	Gen. et sp. nov	Valid	Aristov & Storozhenko	Triassic (Ladinian or Carnian)	Madygen Formation	Kyrgyzstan	A webspinner belonging to the family Alexarasniidae. The type species is M. rasnitsyni.
Magicivena^[409]	Fam. et gen. et 3 sp. nov	Valid	Yang et al.	Early Cretaceous	Yixian Formation	China	A member of Polyneoptera of uncertain phylogenetic placement, belonging to the new family Magicivenidae. The type species is M. sticta; genus also includes M. elegans and M. antennalis.
Multivena^[406]	Gen. et sp. nov	In press	Lai et al.	Cretaceous	Burmese amber	Myanmar	A webspinner belonging to the family Sorellembiidae. Genus includes new species M. curvivena.
Pictoborella^[402]	Gen. et sp. et comb. nov	Valid	Cui et al.	Early Permian		Germany	A member of Grylloblattida. Genus includes new species P. clara, as well as "Oborella" germanica Prokop et al. (2021).
Rasnalexia^[410]	Fam. et gen. et sp. nov	Valid	Gorochov	Triassic		Kyrgyzstan	A member of Embioptera belonging to the new family Rasnalexiidae. Genus includes new species R. rasnitsyni.
Uralioma thuringiensis^[402]	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of Grylloblattida belonging to the family Liomopteridae.

Other insect research

A well-preserved full-body impression of an elongate winged insect strongly resembling stick insects is described from the Permian (Wordian) Pelitic Formation (France) by Logghe et al. (2021), who name a new ichnotaxon Phasmichnus radagasti.^[411]
A dragonfly-type odonatopteran larva, preserving ancestral morphological characters maintained in mayflies but lost in modern odonatan larvae, is described from the Cretaceous Burmese amber by Haug, Müller & Haug (2021).^[412]

General research

Revision of the fossil record of insects from the Triassic of China is published by Zhang et al. (2021).^[413]
A study on the diversity of insect damage types in fossil plants from the Cretaceous (Albian to Cenomanian) Dakota Formation (United States), evaluating their implications for the knowledge of the early evolution of angiosperm florivory and associated pollination, is published by Xiao et al. (2021).^[414]

References

^ ^a ^b Mey, W.; Léger, T.; Lien, V. V. (2021). "New taxa of extant and fossil primitive moths in South-East Asia and their biogeographic significance (Lepidoptera, Micropterigidae, Agathiphagidae, Lophocoronidae)". Nota Lepidopterologica. 44: 29–56. doi:10.3897/nl.44.52350.
^ ^a ^b Fischer, T. C. (2021). "In search for the unlikely: Leaf-mining caterpillars (Gracillariidae, Lepidoptera) from Upper Cretaceous and Eocene ambers". Zitteliana. 95: 135–145. doi:10.3897/zitteliana.95.63317. S2CID 245086588.
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^ Chen, Z.-T. (2021). "Balticonemoura bulbosus gen. et sp. nov., a new stonefly of Nemouridae (Insecta: Plecoptera) from Eocene Baltic amber". Historical Biology. 34 (3): 421–424. doi:10.1080/08912963.2021.1922399. S2CID 235528609.
^ Chen, Z.-T.; Xu, C.-P. (2021). "New perlid stonefly (Insecta: Plecoptera) from mid-Cretaceous Burmese amber". Cretaceous Research. 127: Article 104954. Bibcode:2021CrRes.12704954C. doi:10.1016/j.cretres.2021.104954.
^ Jouault, C.; Legendre, F.; Condamine, F. L.; Nel, A. (2021). "A new stonefly species (Plecoptera: Perlodidae) from Eocene Baltic amber and questions on the wing venation potential for species diagnostic of fossil Plecoptera" (PDF). Palaeoentomology. 4 (3): 243–256. doi:10.11646/palaeoentomology.4.3.12. S2CID 237847715.
^ ^a ^b Aristov, D. S. (2021). "New eoblattid insects (Insecta: Eoblattida) from the Upper Permian of Russia". Paleontological Journal. 55 (1): 72–75. doi:10.1134/S0031030121010020. S2CID 232070926.
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^ ^a ^b ^c ^d ^e Cui, Y.; Brauner, S.; Schneider, J. W.; Béthoux, O. (2021). "Grylloblattidan insects from Sperbersbach and Cabarz (Germany), two new early Permian and insect-rich localities". Journal of Paleontology. 96 (2): 355–374. doi:10.1017/jpa.2021.101. S2CID 243486826.
^ ^a ^b Nel, A.; Poschmann, M. J. (2021). "Early Permian acercarian insects (Miomoptera, Palaeomanteidae) from the Saar-Nahe Basin (Rhineland-Palatinate, SW Germany)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 300 (3): 267–277. doi:10.1127/njgpa/2021/0989. S2CID 236233140.
^ Rasnitsyn, A. P.; Aristov, D. S. (2021). "Review of the family Tococladidae Carpenter (Insecta: Eoblattida) from the Permian of North America and Europe". Paleontological Journal. 55 (2): 151–156. doi:10.1134/S003103012102012X. S2CID 233746323.
^ Falières, E.; Engel, M. S.; Nel, A. (2021). "Earliest occurrence of Embiidae: A new genus from earliest Eocene Oise amber (Insecta: Embiodea)". Comptes Rendus Palevol. 20 (38): 799–805. doi:10.5852/cr-palevol2021v20a38.
^ ^a ^b Lai, Z.; Yang, Q.; Shi, C.; Zhang, W.; Ren, D. (2021). "New webspinners (Insecta: Embioptera) from Upper Cretaceous amber of northern Myanmar". Cretaceous Research. 130: Article 105040. doi:10.1016/j.cretres.2021.105040. S2CID 239202878.
^ ^a ^b ^c Bashkuev, A. S.; Sukatsheva, I. D. (2021). "New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)". Palaeoentomology. 4 (5): 453–461. doi:10.11646/palaeoentomology.4.5.11. ISSN 2624-2826. S2CID 240525412.
^ Aristov, D. S.; Storozhenko, S. Yu. (2021). "A new genus of the family Alexarasniidae (Insecta: Embiida) from the Triassic of Kyrgyzstan". Palaeoentomology. 4 (5): 433–435. doi:10.11646/palaeoentomology.4.5.7. S2CID 240549301.
^ Yang, H.; Shih, C.; Rasnitsyn, A. P.; Ren, D.; Gao, T. (2021). "Early Cretaceous enigmatic insect group showing unique wing venations and antennal sensilla". Papers in Palaeontology. 8. doi:10.1002/spp2.1402. S2CID 244172848.
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^ Xiao, L.; Labandeira, C.; Dilcher, D.; Ren, D. (2021). "Florivory of Early Cretaceous flowers by functionally diverse insects: implications for early angiosperm pollination". Proceedings of the Royal Society B: Biological Sciences. 288 (1953): Article ID 20210320. doi:10.1098/rspb.2021.0320. PMC 8207559. PMID 34132112.

[Acanthocorona-1] Mey, W.; Léger, T.; Lien, V. V. (2021). "New taxa of extant and fossil primitive moths in South-East Asia and their biogeographic significance (Lepidoptera, Micropterigidae, Agathiphagidae, Lophocoronidae)". Nota Lepidopterologica. 44: 29–56. doi:10.3897/nl.44.52350.

[ZittelianaFischer-2] Fischer, T. C. (2021). "In search for the unlikely: Leaf-mining caterpillars (Gracillariidae, Lepidoptera) from Upper Cretaceous and Eocene ambers". Zitteliana. 95: 135–145. doi:10.3897/zitteliana.95.63317. S2CID 245086588.

[3] Haug, J. T.; Haug, C. (2021). "A 100 million-year-old armoured caterpillar supports the early diversification of moths and butterflies". Gondwana Research. 93: 101–105. Bibcode:2021GondR..93..101H. doi:10.1016/j.gr.2021.01.009. S2CID 233548334.

[4] Maccracken, S. A.; Sohn, J.-C.; Miller, I. M.; Labandeira, C. C. (2021). "A new Late Cretaceous leaf mine Leucopteropsa spiralae gen. et sp. nov. (Lepidoptera: Lyonetiidae) represents the first confirmed fossil evidence of the Cemiostominae". Journal of Systematic Palaeontology. 19 (2): 131–144. doi:10.1080/14772019.2021.1881177. S2CID 233194385.

[FEE444-5] Melnitsky, S. I.; Ivanov, V. D. (2021). "Two new species of the genus Archaeopolycentra (Trichoptera: Polycentropodidae) from Cretaceous Taimyr amber". Far Eastern Entomologist. 444: 1–7. doi:10.25221/fee.444.1. ISSN 1026-051X. S2CID 244880131.

[6] Wichard, W.; Neumann, C. (2021). "The polycentropodid genus Cernotina (Insecta, Trichoptera) in Miocene Dominican amber". Fossil Record. 24 (1): 129–133. doi:10.5194/fr-24-129-2021.

[7] Melnitsky, S. I.; Ivanov, V. D.; Perkovsky, E. E. (2021). "A new species of the fossil genus Electrotrichia (Insecta: Trichoptera: Hydroptilidae) from Rovno amber (Zhytomyr region, Olevsk locality)". Palaeoentomology. 4 (5): 421–424. doi:10.11646/palaeoentomology.4.5.4. S2CID 240542805.

[PJ555-8] ^ ^a ^b ^c ^d ^e ^f ^g Sukacheva, I. D.; Vasilenko, D. V. (2021). "Caddisflies (Insecta, Trichoptera) from the upper Mesozoic Chernovskie Kopi locality (Transbaikalia): part 2. Larval cases". Paleontological Journal. 55 (5): 545–551. doi:10.1134/S0031030121050154. S2CID 238640223.

[9] Melnitsky, S. I.; Ivanov, V. D.; Perkovsky, E. E. (2021). "A new species of the genus Holocentropus (Trichoptera: Polycentropodidae) from Rovno amber". Zoosystematica Rossica. 30 (2): 298–302. doi:10.31610/zsr/2021.30.2.298. S2CID 245185514.

[Palaeoentomology44Odontoceridae-10] Wichard, W.; Müller, P.; Xu, C.-P. (2021). "The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber". Palaeoentomology. 4 (4): 331–338. doi:10.11646/palaeoentomology.4.4.8. S2CID 239650542.

[11] Melnitsky, S. I.; Ivanov, V. D.; Perkovsky, E. E. (2021). "A new species of Plectrocnemia (Trichoptera: Polycentropodidae) from Rovno amber". Zootaxa. 5006 (1): 106–109. doi:10.11646/zootaxa.5006.1.14. PMID 34810582. S2CID 239644442.

[12] Sukatsheva, I. D.; Aristov, D. S. (2021). "New data on Cretaceous caddisflies family Vitimotauliidae (Insecta, Trichoptera) from Buryatia (Khasurty locality)". Paleontological Journal. 55 (4): 391–395. doi:10.1134/S0031030121330017. S2CID 237376766.

[WangetalCRWormaldia-13] Wang, J.; Zhang, W.; Shih, C.; Ren, D. (2021). "Two new mid-Cretaceous caddisflies (Trichoptera, Philopotamidae) from amber of Northern Myanmar". Cretaceous Research. 126: Article 104891. Bibcode:2021CrRes.12604891W. doi:10.1016/j.cretres.2021.104891.

[Clusiomitidae-14] Roháček, J.; Hoffeins, C. (2021). "Clusiomitidae, a new family of Eocene fossil Acalyptratae, with revision of Acartophthalmites Hennig and Clusiomites Gen. Nov. (Diptera) †". Insects. 12 (12): Article 1123. doi:10.3390/insects12121123. PMC 8705662. PMID 34940210.

[15] Krzemiński, W.; Kopeć, K.; Soszyńska-Maj, A.; Skibińska, K. (2021). "Aleksiana rasnitsyni gen. et sp. nov. (Diptera, Limoniidae) to honour Alexandr Rasnitsyn". Palaeoentomology. 4 (5): 436–440. doi:10.11646/palaeoentomology.4.5.8. S2CID 240522901.

[Palaeoentomology46Nel-16] Nel, A. (2021). "New Cecidomyiidae from the Lowermost Eocene French amber (Diptera)". Palaeoentomology. 4 (6): 620–628. doi:10.11646/palaeoentomology.4.6.11. S2CID 245179601.

[17] Bouju, V.; Rosse-Guillevic, S.; Griffon, M.; Bojarski, B.; Szwedo, J.; Perrichot, V. (2021). "The genus Allodia (Diptera: Mycetophilidae) in Miocene Ethiopian amber". Fossil Record. 24 (2): 339–346. doi:10.5194/fr-24-339-2021.

[PalaeoworldPhoridae-18] Solórzano-Kraemer, M. M.; Bourdeau, C.; Rosse-Guillevic, S.; Hammel, J. U.; Xu, C.; Brown, B. V. (2021). "Parasitic hump-backed flies (Diptera: Phoridae) from Middle Miocene ambers" (PDF). Palaeoworld. in press. doi:10.1016/j.palwor.2021.11.002. S2CID 244126413.

[19] Lamas, C. J. E.; Sampronha, S.; Ribeiro, G. C. (2021). "A new robber fly from the Lower Cretaceous (Aptian) Crato Formation of NE Brazil (Insecta: Diptera: Asilidae)". Cretaceous Research. 131: Article 105114. doi:10.1016/j.cretres.2021.105114.

[20] Lukashevich, E. D. (2021). "The oldest Diptera (Insecta) from the Upper Buntsandstein (early Middle Triassic) of Europe". Zootaxa. 5067 (1): 135–143. doi:10.11646/zootaxa.5067.1.10. PMID 34810754. S2CID 243901965.

[21] Skartveit, J.; Wedmann, S. (2021). "A revision of fossil Bibionidae (Insecta: Diptera) from the Oligocene of Germany". Zootaxa. 4909 (1): zootaxa.4909.1.1. doi:10.11646/zootaxa.4909.1.1. PMID 33756592. S2CID 232338865.

[22] Skartveit, J. (2021). "A new fossil species of the genus Bibio, with an update on bibionid flies from Baltic and Rovno amber (Diptera, Bibionidae)". Deutsche Entomologische Zeitschrift. 68 (1): 81–99. doi:10.3897/dez.68.60611. hdl:11250/2731857.

[23] Ševčík, J.; Skartveit, J.; Krzemiński, W.; Skibińska, K. (2021). "A Peculiar New Genus of Bibionomorpha (Diptera) with Brachycera-Like Modification of Antennae from Mid-Cretaceous Amber of Myanmar". Insects. 12 (4): Article 364. doi:10.3390/insects12040364. PMC 8073775. PMID 33923888.

[Burmasymmerus-24] Ševčík, J.; Hippa, H.; Burdíková, N. (2021). "Just a Fragment of Undescribed Diversity: Twenty New Oriental and Palearctic Species of Sciaroidea (Diptera), including DNA Sequence Data and Two New Fossil Genera". Insects. 13 (1): Article 19. doi:10.3390/insects13010019. PMC 8779109. PMID 35055862.

[25] Poinar, G.; Vega, F. E. (2021). "A new genus of Apsilocephalidae (Diptera) in Mid-Cretaceous Burmese amber". Biosis: Biological Systems. 2 (1): 183–190. doi:10.37819/biosis.002.01.0091.

[26] Liu, Y.M.; Huang, D.Y. (2021). "A new species of Crenoptychoptera Kalugina, 1985 (Diptera: Ptychopteridae) from the Middle–Late Jurassic of Jiyuan Basin, China". Palaeoentomology. 4 (1): 27–29. doi:10.11646/palaeoentomology.4.1.5. S2CID 233927262.

[27] Lukashevich, E. D.; Amorim, D. de S.; Ribeiro, G. C. (2021). "A Gondwanan record of the extinct genus Cretobibio (Diptera: Bibionidae)". Palaeoentomology. 4 (5): 468–474. doi:10.11646/palaeoentomology.4.5.13. S2CID 244210389.

[SRCretolimonia-28] Kopeć, K.; Soszyńska-Maj, A.; Kania-Kłosok, I.; Coram, R. A.; Krzemiński, W. (2021). "Morphology of the oldest fossil subfamily of Limoniidae (Diptera, Architipulinae) in the light of exceptionally preserved Mesozoic material". Scientific Reports. 11 (1): Article number 24137. Bibcode:2021NatSR..1124137K. doi:10.1038/s41598-021-03350-4. PMC 8683464. PMID 34921169.

[29] Carmo, D. d.D.; Lamas, C. J. E.; Ribeiro, G. C. (2021). "The oldest fossil Stiletto fly: a new genus and species from the Lower Cretaceous Crato Formation of Brazil (Diptera: Therevidae)". Cretaceous Research. 130: Article 105039. doi:10.1016/j.cretres.2021.105039. S2CID 244212150.

[30] Li, L.; Zhang, Q.; Xiao, C. (2021). "A new genus and species of Bibionidae (Diptera: Bibionoidea) from mid-Cretaceous Kachin amber". Acta Palaeontologica Sinica. 60 (2): 243–252. doi:10.19800/j.cnki.aps.2021018.

[31] Giłka, W.; Harbach, R. E.; Perkovsky, E. E. (2021). "Mosquitoes (Diptera: Culicidae) in Eocene amber from the Rovno region, Ukraine". Zootaxa. 5016 (2): 257–270. doi:10.11646/zootaxa.5016.2.6. PMID 34810451. S2CID 238838914.

[HBAnglesea-32] Peñalver, E.; Arillo Aranda, A.; Szadziewski, R.; Stilwell, J. D. (2021). "Biting midges (Diptera: Ceratopogonidae) from the middle Eocene Anglesea amber (Australia) originated in a subpolar greenhouse earth". Historical Biology. 34 (3): 425–435. doi:10.1080/08912963.2021.1924700. S2CID 236275915.

[SRCyttaromyia-33] Kania-Kłosok, I.; Nel, A.; Szwedo, J.; Jordan-Stasiło, W.; Krzemiński, W. (2021). "Phylogeny and biogeography of the enigmatic ghost lineage Cylindrotomidae (Diptera, Nematocera)". Scientific Reports. 11 (1): Article number 13916. Bibcode:2021NatSR..1113916K. doi:10.1038/s41598-021-91719-w. PMC 8260593. PMID 34230501.

[34] Krzemiński, W.; Kania-Kłosok, I.; Krzemińska, E.; Ševčík, J.; Soszyńska-Maj, A. (2021). "Fossils Shed a New Light on the Diversity and Disparity of the Family Limoniidae (Diptera, Nematocera)". Insects. 12 (3): Article 206. doi:10.3390/insects12030206. PMC 8000698. PMID 33804356.

[SRElephantomyia-35] Kania-Kłosok, I.; Krzemiński, W. (2021). "Recent discoveries of new Elephantomyia (Diptera, Limoniidae) fossils in Baltic amber". Scientific Reports. 11 (1): Article number 23647. Bibcode:2021NatSR..1123647K. doi:10.1038/s41598-021-03022-3. PMC 8655017. PMID 34880309.

[36] Szadziewski, R.; Sontag, E.; Pankowski, M. V. (2021). "A new biting midge of the genus Forcipomyia Meigen, 1818 from Miocene Ethiopian amber (Diptera: Ceratopogonidae)". Palaeoentomology. 4 (5): 441–444. doi:10.11646/palaeoentomology.4.5.9. ISSN 2624-2826. S2CID 244172069.

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[Electroxylita-112] ^ ^a ^b ^c ^d ^e ^f Alekseev, V. I.; Bukejs, A. (2021). "Fossil Melandryidae (Coleoptera: Tenebrionoidea) from Eocene Baltic amber of the Sambian peninsula: new genus, six new species, new records and key to described taxa". Zootaxa. 4965 (1): 142–166. doi:10.11646/zootaxa.4965.1.7. PMID 33903499. S2CID 233408753.

[113] Vitali, F. "Eurocrossotus alekseevi gen. n., sp. n. (Coleoptera: Cerambycidae) from Baltic amber, the first fossil member of the tribe Crossotini Thomson, 1864". Baltic Journal of Coleopterology. 21 (2): 221–228.

[114] Legalov, A. A.; Nazarenko, V. Y.; Perkovsky, E. E. (2021). "A new species of the genus Glaesotropis Gratshev and Zherikhin, 1995 (Coleoptera, Anthribidae) from Rovno amber". Fossil Record. 24 (1): 1–7. doi:10.5194/fr-24-1-2021.

[115] Telnov, D.; Perkovsky, E. E.; Vasilenko, D. V.; Yamamoto, S. (2021). "The first fossil Coleoptera record from the Volyn Region, Ukraine, with description of a new Glesoconomorphus (Coleoptera, Mycteridae) in syninclusion with Winterschmidtiidae (Acari) and a key to species". ZooKeys (1068): 189–201. doi:10.3897/zookeys.1068.75391. PMC 8592962. PMID 34819765.

[116] Yu, Y.-L.; Li, Y.-D.; Kolibáč, J.; Ślipiński, A.; Ren, D.; Pang, H.; Li, Z.-Q. (2021). "A New Genus and Species of Lophocateridae from Mid-Cretaceous Amber of Myanmar (Coleoptera)". Insects. 12 (12): Article 1052. doi:10.3390/insects12121052. PMC 8709423. PMID 34940140.

[117] Kupryjanowicz, J.; Lyubarsky, G. Yu.; Perkovsky, E. E. (2021). "Heterhelus buzina sp.n. (Coleoptera: Kateretidae) from Rovno amber: the first proxy for Sambucus in the Eocene of Eastern Europe". Invertebrate Zoology. 18 (1): 16–24. doi:10.15298/invertzool.18.1.02. S2CID 234840368.

[118] Kirejtshuk, A. G.; Jenkins Shaw, J.; Smirnov, I. S. (2023). "A New Subgenus of the Genus Phenolia (Coleoptera, Nitidulidae) from Myanmar Cretaceous Amber with Taxonomic, Phylogenetic and Bionomic Notes on the 'Nitidulid' Group of Families". Insects. 14 (7). 647. doi:10.3390/insects14070647. PMC 10380190. PMID 37504653.

[119] Nabozhenko, M. V.; Bukejs, A. (2021). "A new species and a key to Isomira Mulsant, 1856 (Coleoptera: Tenebrionidae: Alleculinae) from Eocene Baltic amber". Caucasian Entomological Bulletin. 17 (1): 51–56. doi:10.23885/181433262021171-5156.

[120] Legalov, A. A.; Kupryjanowicz, J.; Perkovsky, E. E. (2021). "A new genus of the tribe Cossonini (Coleoptera: Curculionidae) in Baltic amber (Poland)". Paleontological Journal. 55 (4): 405–409. doi:10.1134/S0031030121040109. S2CID 237378812.

[121] Tihelka, E.; Yamamoto, S.; Huang, D.; Cai, C. (2021). "Enigmatic tenebrionoid beetles from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea)". Cretaceous Research. 123: Article 104779. Bibcode:2021CrRes.12304779T. doi:10.1016/j.cretres.2021.104779. S2CID 233908153.

[122] Kirejtshuk, A. G.; Reid, C. A. M. (2022). "Lobanoviella andreyi, new genus and species of Lobanoviellini trib. nov. in the subfamily Palophaginae (Coleoptera, Megalopodidae) from the Eocene Baltic amber, with discussion of similarity with Phloeostichidae". Historical Biology. 34 (9): 1894–1900. doi:10.1080/08912963.2021.1987428. S2CID 243581689.

[Melanosiagon-123] Batelka, J.; Prokop, J. (2021). "The earliest beetle with mouthparts specialized for feeding on nectar is a parasitoid of mid-Cretaceous Hymenoptera". BMC Ecology and Evolution. 21 (1): Article number 207. doi:10.1186/s12862-021-01930-6. PMC 8607574. PMID 34809578.

[124] Nabozhenko, M. V.; McKellar, R. C.; Bukejs, A. (2021). "The first described darkling beetle of the tribe Metaclisini (Coleoptera: Tenebrionidae) from Eocene Baltic amber". Zootaxa. 4999 (3): 279–284. doi:10.11646/zootaxa.4999.3.7. PMID 34810485. S2CID 237770026.

[125] Li, Y.-D.; Tihelka, E.; Leschen, R. A. B.; Yu, Y.; Ślipiński, A.; Pang, H.; Huang, D.; Kolibáč, J.; Cai, C. (2021). "An exquisitely preserved tiny bark-gnawing beetle (Coleoptera: Trogossitidae) from mid-Cretaceous Burmese amber and the phylogeny of Trogossitidae". Journal of Zoological Systematics and Evolutionary Research. 59 (8): 1939–1950. doi:10.1111/jzs.12515. S2CID 238712220.

[126] Bukejs, A.; Bezděk, J.; Kairišs, K.; Alekseev, V. I. (2021). "The first extinct species of Monolepta Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) from Bitterfeld amber, described using X-ray micro-computed tomography". Zootaxa. 4995 (2): 345–356. doi:10.11646/zootaxa.4995.2.7. PMID 34810567. S2CID 237740153.

[127] Zhang, Y. (2021). "New replacement name for the genus Latitergum Yu, Ślipiński, Leschen, Ren & Pang, 2014 (Insecta: Coleoptera: Trogossitidae)". Zootaxa. 4980 (2): 395–396. doi:10.11646/zootaxa.4980.2.11. PMID 34186972. S2CID 235687049.

[128] Szawaryn, K.; Sontag, E.; Kubisz, D. (2021). "The first described fossil Oedemeridae (Insecta: Coleoptera) from Baltic amber". Journal of Paleontology. 96 (2): 375–378. doi:10.1017/jpa.2021.102. S2CID 242027889.

[129] Tihelka, E.; Huang, D.; Cai, C. (2021). "A new genus of giant sooty mould beetles from mid-Cretaceous Burmese amber (Coleoptera: Cyclaxyridae)". Cretaceous Research. 125: Article 104876. Bibcode:2021CrRes.12504876T. doi:10.1016/j.cretres.2021.104876.

[130] Legalov, A. A. (2021). "First record of the subfamily Sagrinae (Coleoptera: Chrysomelidae) from the Eocene of North America". Fossil Record. 24 (1): 135–139. doi:10.5194/fr-24-135-2021.

[131] Legalov, A.A. (2021). "New tribe Palaeoanoplini trib. nov. (Coleoptera: Curculionidae) in Baltic amber". Paleontological Journal. 55 (2): 179–183. doi:10.1134/S0031030121020076. ISSN 0031-0301. S2CID 233746782.

[132] Poinar, G. O.; Brown, A. E.; Legalov, A. A. (2021). "First record of the subfamily Chilecarinae (Coleoptera; Ithyceridae) from mid-Cretaceous Burmese amber". Cretaceous Research. 124: Article 104793. Bibcode:2021CrRes.12404793P. doi:10.1016/j.cretres.2021.104793. S2CID 233806293.

[133] Powell, G. S.; Cline, A. R. (2021). "The First Cillaeinae (Coleoptera: Nitidulidae) Described from Amber". Annales Zoologici. 71 (1): 21–26. doi:10.3161/00034541ANZ2021.71.1.003. S2CID 232422254.

[134] Tihelka, E.; Li, L.; Fu, Y.; Su, Y.; Huang, D.; Cai, C. (2021). "Angiosperm pollinivory in a Cretaceous beetle". Nature Plants. 7 (4): 445–451. doi:10.1038/s41477-021-00893-2. PMID 33846595. S2CID 233223519.

[135] Bao, T.; Wedmann, S.; Grímsson, F.; Beutel, R. G.; Seyfullah, L.; Bao, L.; Jarzembowski, E. A. (2021). "Was the kateretid beetle Pelretes really a Cretaceous angiosperm pollinator?". Nature Plants. 8 (1): 38–40. doi:10.1038/s41477-021-01044-3. PMID 34949805. S2CID 245471963.

[136] Tihelka, E.; Li, L.; Fu, Y.; Su, Y.; Huang, D.; Cai, C. (2021). "Reply to: Pelretes vivificus was a pollinator of Cretaceous angiosperms". Nature Plants. 8 (1): 41–44. doi:10.1038/s41477-021-01045-2. PMID 34949806. S2CID 245445815.

[137] Peris, D.; Jelínek, J.; Sabatelli, S.; Liu, M.-K.; Peña-Kairath, C.; Zhao, Q.; Cai, C.-Y.; Kairišs, K.; Mähler, B.; Rühr, P. T.; Hammel, J. U.; Audisio, P. (2024). "Archaic sap beetles (Coleoptera: Nitidulidae) as Cretaceous pollinators". Palaeoentomology. 7 (5): 594–610. doi:10.11646/palaeoentomology.7.5.4.

[138] Kirejtshuk, A. G.; Jenkins Shaw, J.; Smirnov, I. S. (2023). "A New Subgenus of the Genus Phenolia (Coleoptera, Nitidulidae) from Myanmar Cretaceous Amber with Taxonomic, Phylogenetic and Bionomic Notes on the 'Nitidulid' Group of Families". Insects. 14 (7). 647. doi:10.3390/insects14070647. PMC 10380190. PMID 37504653.

[139] Tihelka, E.; Huang, D.; Perrichot, V.; Cai, C. (2021). "A previously missing link in the evolution of dasytine soft-winged flower beetles from Cretaceous Charentese amber (Coleoptera, Melyridae)" (PDF). Papers in Palaeontology. 7 (4): 1753–1764. doi:10.1002/spp2.1360. S2CID 234862770.

[140] Tshernyshev, S.E.; Shcherbakov, M.V. (2021). "A new genus and species of malachite beetles (Coleoptera, Dasytidae) in Baltic amber". Paleontological Journal. 55 (2): 174–178. doi:10.1134/S0031030121020039. S2CID 233746774.

[141] Tshernyshev, S. E.; Perkovsky, E. E. (2021). "Protomauroania mikhailovi—a new species of malachite beetles (Coleoptera, Dasytidae) in Rovno amber". Zootaxa. 5006 (1): 189–194. doi:10.11646/zootaxa.5006.1.20. PMID 34810576. S2CID 243574205.

[142] Bukejs, A.; Ślipiński, A.; Mitchell, J. L.; McKellar, R. C.; Barbi, M.; Larsson, H. C. E.; Alekseev, V. I. (2021). "First fossil representative of Cerylonidae (Coleoptera: Coccinelloidea) described using X-ray micro-computed tomography, from Eocene Baltic amber". Zootaxa. 5032 (2): 225–236. doi:10.11646/zootaxa.5032.2.4. PMID 34811132. S2CID 239630223.

[143] Krell, F.-T.; Vitali, F. (2021). "Attenborough's beauty: exceptional pattern preservation in a frog-legged leaf beetle from the Eocene Green River Formation, Colorado (Coleoptera, Chrysomelidae, Sagrinae)". Papers in Palaeontology. 7 (4): 2101–2112. doi:10.1002/spp2.1398. S2CID 238854571.

[144] Poinar, G.; Brown, A. E. (2021). "A new tribe, genus and species of weevil, Rhamphophorus legalovii gen. et sp. nov., (Coleoptera, Nemonychidae, Rhamphophorini tribe nov.) in Burmese amber". Cretaceous Research. 127: Article 104948. doi:10.1016/j.cretres.2021.104948.

[145] Alekseev, V. I.; Bukejs, A.; Pankowski, M. G.; Ślipiński, A. (2021). "The first representative of the family Teredidae (Coleoptera: Coccinelloidea) in the fossil record". Historical Biology. 34 (11): 1–6. doi:10.1080/08912963.2021.2009474. S2CID 244917429.

[Zherikhiniellus-146] Legalov, A. A. (2021). "New weevils of the family Brentidae (Coleoptera) in Baltic amber". Paleontological Journal. 55 (1): 96–100. doi:10.1134/S003103012101007X. S2CID 232071010.

[147] Li, Y.-D.; Tihelka, E.; Liu, Z.-H.; Huang, D.-Y.; Cai, C.-Y. (2021). "New mid-Cretaceous cryptic slime mold beetles and the early evolution of Sphindidae (Coleoptera: Cucujoidea)". Arthropod Systematics & Phylogeny. 79: 587–597. doi:10.3897/asp.79.e72724.

[148] Lyubarsky, G. Yu.; Tihelka, E.; Cai, C.-Y.; Perkovsky, E. E. (2021). "Zaiwa: A new genus of the family Lophocateridae from mid-Cretaceous Burmese amber (Coleoptera: Cleroidea)". Palaeoentomology. 4 (3): 218–222. doi:10.11646/palaeoentomology.4.3.7. ISSN 2624-2826. S2CID 237862213.

[149] Batelka, J.; Engel, M. S.; Prokop, J. (2021). "The complete life cycle of a Cretaceous beetle parasitoid". Current Biology. 31 (3): R118–R119. doi:10.1016/j.cub.2020.12.007. PMID 33561406. S2CID 231848712.

[150] Poinar, G. O.; Vega, F. E.; Legalov, A. A. (2021). "A New Species of the Genus Amberophytum Yu, Slipinski et Pang, 2019 (Coleoptera: Cerophytidae) from mid-Cretaceous Burmese Amber". Biosis: Biological Systems. 2 (3): 337–341. doi:10.37819/biosis.002.03.0125. S2CID 238697106.

[151] Li, Y.-D.; Kundrata, R.; Packova, G.; Huang, D.-Y.; Cai, C.-Y. (2021). "An unusual elateroid lineage from mid-Cretaceous Burmese amber (Coleoptera: Elateroidea)". Scientific Reports. 11 (1): Article number 21985. Bibcode:2021NatSR..1121985L. doi:10.1038/s41598-021-01398-w. PMC 8578672. PMID 34753998.

[152] Fanti, F. "A new genus of Cantharidae from Eocene Baltic amber found in Poland, with phoretic mites". Baltic Journal of Coleopterology. 21 (2): 105–110.

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[154] Kundrata, R.; Gimmel, M. L.; Packova, G.; Bukejs, A.; Blank, S. M. (2021). "A new enigmatic lineage of Dascillidae (Coleoptera: Elateriformia) from Eocene Baltic amber described using X-ray microtomography, with notes on Karumiinae morphology and classification". Fossil Record. 24 (1): 141–149. doi:10.5194/fr-24-141-2021.

[155] Jiang, R.; Song, H.; Zhang, H.; Wang, S. (2021). "Burmagrilus cretacus gen. et sp. nov., the first Buprestidae from mid-Cretaceous Burmese amber". Cretaceous Research. 125: Article 104866. Bibcode:2021CrRes.12504866J. doi:10.1016/j.cretres.2021.104866.

[CRBurmomiles-156] Yang, Y.; Zhao, W.; Geiser, M.; Zhang, W.; Bai, M.; Liu, H. (2021). "Review of the genus Burmomiles Fanti, Damgaard & Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae)". Cretaceous Research. 128: Article 104961. Bibcode:2021CrRes.12804961Y. doi:10.1016/j.cretres.2021.104961.

[BJC222-157] Fanti, F.; Müller, P. (2022). "Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information". Baltic Journal of Coleopterology. 22 (2): 331–380.

[Captopus-158] Li, Y.; Huang, D.; Cai, C. (2021). "New genera and species of the family Throscidae (Coleoptera: Elateroidea) in mid-Cretaceous Burmese amber". Insects. 12 (1): Article 63. doi:10.3390/insects12010063. PMC 7826609. PMID 33445628.

[FR241Chelonarium-159] Alekseev, V. I.; Mitchell, J.; McKellar, R. C.; Barbi, M.; Larsson, H. C. E.; Bukejs, A. (2021). "The first described turtle beetles from Eocene Baltic amber, with notes on fossil Chelonariidae (Coleoptera: Byrrhoidea)". Fossil Record. 24 (1): 19–32. doi:10.5194/fr-24-19-2021.

[160] Poinar, G.; Háva, J. (2021). "The first representative of the beetle family Chelonariidae in Dominican amber". Palaeodiversity. 14 (1): 1–4. doi:10.18476/pale.v14.a1.

[Cretocantharis-161] Hsiao, Y.; Li, Y.; Ren, D.; Pang, H. (2021). "Morphological phylogenetics provide new insights into the classification and evolution of fossil soldier beetles from Mid-Cretaceous Burmese amber (Coleoptera: Cantharidae)". Zoological Journal of the Linnean Society. 193 (4): 1271–1293. doi:10.1093/zoolinnean/zlaa184.

[162] Kundrata, R.; Prosvirov, A. S.; Long, R.; Packova, G. (2021). "Cretopachyderes gen. nov., a new remarkable click beetle (Coleoptera: Elateridae: Agrypninae) from the mid-Cretaceous Burmese amber". Fossil Record. 24 (2): 347–355. doi:10.5194/fr-24-347-2021.

[163] Li, Y.; Kundrata, R.; Tihelka, E.; Liu, Z.; Huang, D.; Cai, C. (2021). "Cretophengodidae, a new Cretaceous beetle family, sheds light on the evolution of bioluminescence". Proceedings of the Royal Society B: Biological Sciences. 288 (1943): Article ID 20202730. doi:10.1098/rspb.2020.2730. PMC 7893276. PMID 33468008.

[REJ302-164] Kazantsev, S. V. (2021). "New species of Baltic amber soldier beetles (Insecta: Coleoptera: Cantharidae)". Russian Entomological Journal. 30 (2): 153–158. doi:10.15298/rusentj.30.2.08. ISSN 0132-8069. S2CID 242185955.

[BJC211-165] Fanti, F. "Two new Malthodes and a new Podistra from Eocene Baltic amber (Coleoptera, Cantharidae)". Baltic Journal of Coleopterology. 21 (1): 1–9.

[166] Pintilioaie, A.-M.; Fanti, F.; Ionesi, V. (2021). "First finding of a fossilized Cantharidae (Insecta: Coleoptera) from the Middle Miocene deposits in Romania". Travaux du Muséum National d'Histoire Naturelle "Grigore Antipa". 64 (1): 73–80. doi:10.3897/travaux.64.e64706. S2CID 237805010.

[167] Li, Y.; Tihelka, E.; Huang, D.; Cai, C. (2021). "Murcybolus gen. nov., a new net-winged beetle genus from mid-Cretaceous Burmese amber (Coleoptera: Lycidae: Burmolycini)". Zootaxa. 4966 (1): 77–83. doi:10.11646/zootaxa.4966.1.8. PMID 34186632. S2CID 235577673.

[168] Li, Y.-D.; Huang, D.; Cai, C. (2021). "Pseudomataeopsephus, a new genus of water penny beetles from mid-Cretaceous Burmese amber (Coleoptera: Psephenidae)". Cretaceous Research. 130: Article 105055. doi:10.1016/j.cretres.2021.105055. S2CID 244242362.

[169] Kundrata, R.; Packova, G.; Kairišs, K.; Bukejs, A.; Hoffmannova, J.; Blank, S. M. (2021). "The First Ptilodactyla Illiger, 1807 (Coleoptera: Dryopoidea: Ptilodactylidae) Described from Eocene Baltic Amber". Biology. 10 (9): Article 877. doi:10.3390/biology10090877. PMC 8465962. PMID 34571754.

[170] Li, H.; Chang, H.; Muona, J.; Zhao, Y.; Ren, D. (2021). "Subfamily Anischiinae (Coleoptera: Eucnemidae) in Early Cretaceous of Northeast China". Insects. 12 (2): Article 105. doi:10.3390/insects12020105. PMC 7910996. PMID 33530615.

[Zootaxa49961-171] Fanti, F.; Pankowski, M. G. (2021). "Two new species of soldier beetles (Coleoptera, Cantharidae), the first from the tribe Silini in Dominican amber". Zootaxa. 4996 (1): 163–170. doi:10.11646/zootaxa.4996.1.7. PMID 34810540. S2CID 237823176.

[172] Li, Y.-D.; Kundrata, R.; Huang, D.-Y.; Cai, C.-Y. (2021). "Notes on the Morphology and Systematic Position of Archaeolus Lin, 1986, from the Jurassic of South China (Coleoptera: Elateroidea)". Insects. 12 (10): Article 876. doi:10.3390/insects12100876. PMC 8537516. PMID 34680645.

[173] Haug, J. T.; Zippel, A.; Haug, G. T.; Hoffeins, C.; Hoffeins, H.-W.; Hammel, J. U.; Baranov, V.; Haug, C. (2021). "Texas beetle larvae (Brachypsectridae) – the last 100 million years reviewed". Palaeodiversity. 14 (1): 161–183. doi:10.18476/pale.v14.a8.

[174] Matalin, A. V.; Perkovsky, E. E.; Vasilenko, D. V. (2021). "First record of tiger beetles (Coleoptera, Cicindelidae) from Rovno amber, with the description of a new genus and species". Zootaxa. 5016 (2): 243–256. doi:10.11646/zootaxa.5016.2.5. PMID 34810452. S2CID 238845128.

[175] Tello, F.; Verdú, J. R.; Rossini, M.; Zunino, M. (2021). "Onthophagus pilauco sp. nov. (Coleoptera, Scarabaeidae): evidence of beetle extinction in the Pleistocene–Holocene transition in Chilean Northern Patagonia". ZooKeys (1043): 133–145. doi:10.3897/zookeys.1043.61706. PMC 8219658. PMID 34168516.

[176] Alekseev, V. I.; Bukejs, A. (2021). "The first extinct species of Acritus LeConte, 1853 (Histeridae: Abraeinae) from Eocene Baltic amber: a microscopic beetle inclusion studied with X-ray micro-computed tomography". Fossil Record. 24 (2): 223–231. doi:10.5194/fr-24-223-2021.

[177] Alekseev, V. I.; Bukejs, A. (2021). "A new Eocene Bacanius species (Histeridae: Dendrophilinae) from Baltic amber". Fossil Record. 24 (1): 93–99. doi:10.5194/fr-24-93-2021.

[178] Yin, Z.-W.; Cai, C.-Y. (2021). "A new Brachyglutini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Staphylinidae: Pselaphinae)". Cretaceous Research. 124: Article 104807. Bibcode:2021CrRes.12404807Y. doi:10.1016/j.cretres.2021.104807. S2CID 233577289.

[Carinumerus-179] Caterino, M. S. (2021). "New Fossil Histerid Species from Cretaceous Burmese Amber (Coleoptera: Histeridae)". The Coleopterists Bulletin. 75 (1): 211–221. doi:10.1649/0010-065X-75.1.211. S2CID 232368396.

[180] Yamamoto, S.; Shavrin, A. V.; Kairišs, K. (2021). "A second fossil species of the enigmatic rove beetle genus Charhyphus in Eocene Baltic amber, with implications on the morphology of the female genitalia (Coleoptera: Staphylinidae: Phloeocharinae)". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 113: 39–50. doi:10.1017/S1755691021000360. hdl:2115/86861. S2CID 244931145.

[181] Sohn, J.-C.; Nam, G. S. (2021). "New fossil genus and species of carrion beetle (Coleoptera, Silphidae) from the Lower Cretaceous Jinju Formation, South Korea". Journal of Asia-Pacific Entomology. 24 (3): 584–587. doi:10.1016/j.aspen.2021.05.003. S2CID 236585083.

[182] Jałoszyński, P.; Perkovsky, E. E. (2021). "A new bizarre species of Euconnus (Cladoconnus) in Upper Eocene Rovno amber (Coleoptera: Staphylinidae: Scydmaeninae)". Zootaxa. 5004 (2): 395–400. doi:10.11646/zootaxa.5004.2.8. PMID 34811301. S2CID 237709269.

[183] Shavrin, A. V.; Kairišs, K. (2021). "A new species of Eusphalerum Kraatz, 1857 from the Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)". Zootaxa. 4966 (4): 469–475. doi:10.11646/zootaxa.4966.4.5. PMID 34186602. S2CID 235565793.

[184] Yin, Z.-W.; Zhao, Q.-H.; Cai, C.-Y. (2021). "Megabythinus gen. nov., a new stem-group Pselaphinae from mid-Cretaceous Myanmar amber (Coleoptera, Staphylinidae)". Cretaceous Research. 125: Article 104879. Bibcode:2021CrRes.12504879Y. doi:10.1016/j.cretres.2021.104879.

[185] Liu, Y.; Tihelka, E.; Thayer, M. K.; Newton, A. F.; Huang, D.; Tian, L.; Cai, C. (2021). "A transitional fossil sheds light on the early evolution of the Staphylinine group of rove beetles (Coleoptera: Staphylinidae)". Journal of Systematic Palaeontology. 19 (4): 321–332. doi:10.1080/14772019.2021.1917705. S2CID 235172619.

[186] Prokin, A. A.; Strelnikova, O. D. (2021). "The first remarkable fossil Hydrochidae (Coleoptera) from the Lower Cretaceous (Hauterivian) of the Buryatia Republic, Russia". Cretaceous Research. 123: Article 104795. Bibcode:2021CrRes.12304795P. doi:10.1016/j.cretres.2021.104795. S2CID 233957495.

[187] Perreau, M.; Haelewaters, D.; Tafforeau, P. (2021). "A parasitic coevolution since the Miocene revealed by phase-contrast synchrotron X-ray microtomography and the study of natural history collections". Scientific Reports. 11 (1): Article number 2672. Bibcode:2021NatSR..11.2672P. doi:10.1038/s41598-020-79481-x. PMC 7846571. PMID 33514784.

[188] Yamamoto, S.; Newton, A. F. (2021). "First fossil trichophyine rove beetle from mid-Cretaceous amber of northern Myanmar (Coleoptera: Staphylinidae: Trichophyinae)". Cretaceous Research. 127: Article 104951. Bibcode:2021CrRes.12704951Y. doi:10.1016/j.cretres.2021.104951. hdl:2115/90159.

[189] Zhao, X.; Yu, Y.; Clapham, M. E.; Yan, E.; Chen, J.; Jarzembowski, E. A.; Zhao, X.; Wang, B. (2021). "Early evolution of beetles regulated by the end-Permian deforestation". eLife. 10: e72692. doi:10.7554/eLife.72692. PMC 8585485. PMID 34747694.

[190] Li, Y.-D.; Huang, D.-Y.; Cai, C.-Y. (2021). "Revisiting the systematic positions of two "Notocupes" species from the Lower Cretaceous of South China (Coleoptera: Polyphaga)". Zootaxa. 5005 (1): 86–90. doi:10.11646/zootaxa.5005.1.7. PMID 34811277. S2CID 237669437.

[191] Haug, C.; Haug, G. T.; Zippel, A.; van der Wal, S.; Haug, J. T. (2021). "The earliest record of fossil solid-wood-borer larvae—immature beetles in 99 million-year-old Myanmar amber". Palaeoentomology. 4 (4): 390–404. doi:10.11646/palaeoentomology.4.4.14. ISSN 2624-2826. S2CID 237622518.

[192] Smith, A. D.; Kamiński, M. J.; Kanda, K.; Sweet, A. D.; Betancourt, J. L.; Holmgren, C. A.; Hempel, E.; Alberti, F.; Hofreiter, M. (2021). "Recovery and analysis of ancient beetle DNA from subfossil packrat middens using high-throughput sequencing". Scientific Reports. 11 (1): Article number 12635. doi:10.1038/s41598-021-91896-8. PMC 8209150. PMID 34135378.

[Afrophaga-193] ^ ^a ^b ^c ^d ^e ^f ^g Vršanský, P.; Vršanská, L.; Vasilenko, D. V.; Puškelová, L.; Biroň, A. (2021). "An isolated Cretaceous analogue of Madagascar on the Adria–Turkey microcontinent indicated by fossils in Brezina, Algeria". Palaeontographica Abteilung A. 321 (1–6): 19–35. doi:10.1127/pala/2021/0107. S2CID 240038689.

[Nadveruzenie-194] Vršanský, P.; Sendi, H.; Hinkelman, J.; Hain, M. (2021). "Alienopterix Mlynský et al., 2018 complex in North Myanmar amber supports Umenocoleoidea/ae status". Biologia. 76 (8): 2207–2224. doi:10.1007/s11756-021-00689-x. S2CID 233784192.

[Arceotermes-195] Jiang, R.X.; Zhang, H.R.; Eldredge, K.T.; Song, X.B.; Li, Y.D.; Tihelka, E.; Huang, D.Y.; Wang, S.; Engel, M.S.; Cai, C.Y. (2021). "Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)". Palaeoentomology. 4 (4): 374–389. doi:10.11646/palaeoentomology.4.4.13. S2CID 239695173.

[Caligoptera-196] Sendi, H. (2021). "Highly specialised basal ectobiid cockroaches (Blattaria: Blattoidea) were rare in Burmese amber". Palaeontographica Abteilung A. 321 (1–6): 109–125. doi:10.1127/pala/2021/0106. S2CID 243471520.

[197] Liang, J.; Wang, Y.; Shih, C.; Ren, D. (2021). "Chuanblatta gen. nov. sexually dimorphic cockroaches of Raphidiomimidae (Blattaria) from the Jiulongshan Formation in China". Palaeontographica Abteilung A. 321 (1–6): 3–17. doi:10.1127/pala/2021/0113. ISSN 0375-0442. S2CID 240180893.

[198] Hinkelman, J. (2021). "Cuniculoblatta brevialata gen. et sp. n., the second case of brachyptery from Cretaceous North Myanmar amber". Palaeontographica Abteilung A. 321 (1–6): 97–107. doi:10.1127/pala/2021/0104. S2CID 240183883.

[199] Chen, X.-Y.; Zhang, H.-C.; Shi, X. (2021). "A new species and genus of Alienopteridae (Blattodea) from mid-Cretaceous amber of northern Myanmar". Zootaxa. 4941 (4): 580–586. doi:10.11646/zootaxa.4941.4.7. PMID 33756925. S2CID 232339877.

[Fractalia-200] Vršanský, P.; Hinkelman, J.; Koubová, I.; Sendi, H.; Kúdelová, T.; Kúdela, M.; Barclay, M. (2021). "A single common ancestor for praying mantids, termites, cave roaches and umenocoleoids" (PDF). AMBA Projekty. 11 (1): 1–16.

[201] Chen, G.; Xiao, L.; Liang, J.; Shih, C.; Ren, D. (2021). "A new cockroach (Blattodea, Corydiidae) with pectinate antennae from mid-Cretaceous Burmese amber". ZooKeys (1060): 155–169. doi:10.3897/zookeys.1060.67216. PMC 8486729. PMID 34690506.

[202] Bouju, V.; Jouault, C.; Perrichot, V. (2021). "The termite genus Glyptotermes (Isoptera: Kalotermitidae) in Miocene amber from Ethiopia" (PDF). Journal of Paleontology. 96 (2): 387–393. doi:10.1017/jpa.2021.106. S2CID 244426915.

[203] Schneider, J. W.; Scholze, F.; Germann, S.; Lucas, S. G. (2021). "The Late Pennsylvanian nearshore insect fauna of the Kinney Brick Quarry invertebrate and vertebrate fossil Lagerstätte, New Mexico". New Mexico Museum of Natural History and Science Bulletin. 84: 255–285.

[204] Demers-Potvin, A. V.; Larsson, H. C. E.; Cournoyer, M.; Béthoux, O. (2021). "Wing morphology of a new Cretaceous praying mantis solves the phylogenetic jigsaw of early-diverging extant lineages". Systematic Entomology. 46 (1): 205–223. doi:10.1111/syen.12457. S2CID 231391397.

[205] Vršanský, P.; Poschmann, M. J.; Vidlička, L. (2021). "Oligocene pseudophyllodromiin cockroach from the Enspel Fossillagerstätte in Germany". Palaeontographica Abteilung A. 321 (1–6): 149–167. doi:10.1127/pala/2021/0110. ISSN 0375-0442. S2CID 240039709.

[206] Luo, C.; Beutel, R. G.; Xu, C.; Jarzembowski, E. A. (2021). "†Laticephalana liuyani gen. et sp. nov., a new bizarre roachoid of †Umenocoleidae (Insecta, Dictyoptera) from mid-Cretaceous Kachin amber". Proceedings of the Geologists' Association. 132 (4): 469–478. Bibcode:2021PrGA..132..469L. doi:10.1016/j.pgeola.2021.04.004. S2CID 236329819.

[207] Perkovsky, E. E.; Nel, A. (2021). "A new Rovno amber termite genus (Isoptera, Rhinotermitidae) from Styr river basin". Palaeontologia Electronica. 24 (1): Article number 24.1.a05. doi:10.26879/1127.

[208] Bezerra, F. I.; DeSouza, O.; Ribeiro, G. C.; Mendes, M. (2021). "A new primitive termite (Isoptera) from the Crato Formation, Araripe Basin, Early Cretaceous of South America". Journal of South American Earth Sciences. 109: Article 103260. Bibcode:2021JSAES.10903260B. doi:10.1016/j.jsames.2021.103260. S2CID 233807443.

[Milesitermes-209] Jouault, C.; Legendre, F.; Grandcolas, P.; Nel, A. (2021). "Revising dating estimates and the antiquity of eusociality in termites using the fossilized birth–death process". Systematic Entomology. 46 (3): 592–610. doi:10.1111/syen.12477. S2CID 235363027.

[Miniblattina-210] Sendi, H. (2021). "Diverse Liberiblattinidae (Insecta: Blattaria) from Lebanese and North Myanmar amber document allometric modifications near lowest size limit". Palaeontographica Abteilung A. 321 (1–6): 127–148. doi:10.1127/pala/2021/0108. S2CID 243475716.

[211] Hinkelman, J. (2021). "Mongolblatta sendii sp. n. (Mesoblattinidae) from North Myanmar amber links record to Laurasian sediments". Palaeontographica Abteilung A. 321 (1–6): 81–96. doi:10.1127/pala/2021/0105. S2CID 240162578.

[PAABlaberidae-212] Šmídová, L.; Vidlička, L.; Wedmann, S. (2021). "Appearance of the family Blaberidae (Insecta: Blattaria) during the Cretaceous and a review of fossils of this family". Palaeontographica Abteilung A. 321 (1–6): 71–79. doi:10.1127/pala/2021/0109. S2CID 243467634.

[Olenablatta-213] Šmídová, L. (2021). "New genus and species of the families Olidae and Corydiidae (Corydioidea, Blattodea) from mid-Cretaceous Kachin amber". Palaeontographica Abteilung A. 321 (1–6): 61–70. doi:10.1127/pala/2021/0117. S2CID 243471507.

[214] Song, Z.; Xu, C.; Li, J.; Jarzembowski, E. A.; Wang, B.; Xiao, C. (2021). "A new species of Pabuonqedidae (Blattaria: Mastotermitoidea) from mid-Cretaceous Kachin amber". Palaeontographica Abteilung A. 321 (1–6): 53–59. doi:10.1127/pala/2021/0111. S2CID 240023055.

[215] So, K. S.; Won, C. G.; Ri, C. J.; Jon, S. H.; Ju, I. Y. (2021). "Paekthoblatta, a New Predatory Cockroach Genus (Insecta: Blattaria: Raphidiomimidae) from the Lower Cretaceous of Paektho-Dong, Sinuiju, Democratic People's Republic of Korea". Paleontological Journal. 55 (8): 906–909. doi:10.1134/S0031030121080074. S2CID 245330342.

[216] Anisyutkin, L. N.; Perkovsky, E. E. (2021). "Periplaneta (?) perialla sp. nov., a cockroach larva of the subfamily Blattinae (Dictyoptera, Blattidae) from Rovno amber". Paleontological Journal. 55 (3): 294–300. doi:10.1134/S0031030121030047. S2CID 235966737.

[OyamaetalPAA-217] Oyama, N.; Yukawa, H.; Imai, T. (2021). "New cockroach assemblage from the Lower Cretaceous Kitadani Formation, Fukui, Japan". Palaeontographica Abteilung A. 321 (1–6): 37–52. doi:10.1127/pala/2021/0112. S2CID 239140974.

[218] Jiang, L.; Xing, J.; Li, Y. (2023). "New replacement name for the genus Sajda Vršanský, 2021 (Blattaria: Corydiidae: Holocompsinae)". Zootaxa. 5270 (2): 343–344. doi:10.11646/zootaxa.5270.2.10. PMID 37518160. S2CID 258274946.

[219] So, K. S.; Won, C. G.; Ri, C. J.; Jon, S. H.; Ju, I. Y. (2021). "A New Species of Spinaeblattina Hinkelman, 2019 (Insecta, Blattaria, Mesoblattinidae) from the Lower Cretaceous of Paektho-Dong, Sinuiju, Democratic People's Republic of Korea". Paleontological Journal. 55 (8): 910–912. doi:10.1134/S0031030121080086. S2CID 245330596.

[220] Liuo, C.-H.; Beutel, R. G.; Thomson, U. R; Zheng, D.-R.; Li, J.-H.; Zhao, X.-Y.; Zhang, H.-C.; Wang, B. (2021). "Beetle or roach: systematic position of the enigmatic Umenocoleidae based on new material from Zhonggou Formation in Jiuquan, Northwest China, and a morphocladistic analysis". Palaeoworld. 31: 121–130. doi:10.1016/j.palwor.2021.01.003. S2CID 234208413.

[221] Luo, C.; Beutel, R. G.; Engel, M. S.; Liang, K.; Li, L.; Li, J.; Xu, C.; Vršanský, P.; Jarzembowski, E. A.; Wang, B. (2021). "Life history and evolution of the enigmatic Cretaceous-Eocene Alienopteridae: A critical review". Earth-Science Reviews. 225: Article 103914. doi:10.1016/j.earscirev.2021.103914. S2CID 245601183.

[222] Taniguchi, R.; Nishino, H.; Watanabe, H.; Yamamoto, S.; Iba, Y. (2021). "Reconstructing the ecology of a Cretaceous cockroach: destructive and high-resolution imaging of its micro sensory organs". The Science of Nature. 108 (5): Article number 45. Bibcode:2021SciNa.108...45T. doi:10.1007/s00114-021-01755-9. ISSN 0028-1042. PMID 34581877. S2CID 238202024.

[Moltenojurina-223] Béthoux, O.; Anderson, J. M. (2021). "The Polycytellidae viewed as Gondwanan Glosselytrodea". Palaeoentomology. 4 (6): 550–558. doi:10.11646/palaeoentomology.4.6.5. S2CID 245173292.

[Aethemelikertes-224] Engel, M. S.; Davis, S. R. (2021). "New genera of melikertine bees with facial modifications in Baltic amber (Hymenoptera: Apidae)". Journal of Melittology. 103 (103): 1–52. doi:10.17161/jom.i103.15655. S2CID 237834516.

[CRVespidae-225] Wu, Q.; Yang, Y.; Shih, C.; Zhao, Y.; Ren, D. (2021). "Two new species of Vespidae (Hymenoptera) in mid-Cretaceous amber from northern Myanmar". Cretaceous Research. 132: Article 105117. doi:10.1016/j.cretres.2021.105117.

[Allommation-226] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o Rosa, B. B.; Melo, G. A. R. (2021). "Apoid wasps (Hymenoptera: Apoidea) from mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 122: Article 104770. Bibcode:2021CrRes.12204770R. doi:10.1016/j.cretres.2021.104770. S2CID 234071940.

[227] Ngô-Muller, V.; Garrouste, R.; Schubnel, T.; Pouillon, J.-M.; Christophersen, V.; Christophersen, N.; Nel, A. (2021). "The first representative of the trap-jaw ant genus Anochetus Mayr, 1861 in Neogene amber from Sumatra (Hymenoptera: Formicidae)". Comptes Rendus Palevol. 20 (2): 21–27. doi:10.5852/cr-palevol2021v20a2.

[228] Tribull, C. M.; Pankowski, M. V.; Colombo, W. D. (2021). "A new genus and species of Pristocerinae (Hymenoptera, Bethylidae) from upper Eocene Baltic amber with a review of conspecific association from insect fossils". Journal of Hymenoptera Research. 85: 119–133. doi:10.3897/jhr.85.68658.

[229] Rasnitsyn, A. P. (2021). "First Jurassic representative of the extinct family Peleserphidae (Hymenoptera, Proctotrupoidea)". Journal of Hymenoptera Research. 84: 295–300. doi:10.3897/jhr.84.65493.

[Palaeocolastes-230] Belokobylskij, S. A.; Pankowski, M. G.; Pankowski, M. V.; Zaldívar-Riverón, A. (2021). "A new genus and three new species of fossil braconid wasps (Hymenoptera, Ichneumonoidea) from Eocene Baltic and Rovno ambers". Journal of Paleontology. 95 (6): 1259–1272. Bibcode:2021JPal...95.1259B. doi:10.1017/jpa.2021.47. S2CID 237921761.

[JM105-231] Engel, M. S.; Herhold, H.; Davis, S.; Wang, B.; Thomas, J. (2021). "Stingless bees in Miocene amber of southeastern China (Hymenoptera: Apidae)". Journal of Melittology. 105 (105): 1–83. doi:10.17161/jom.i105.15734. S2CID 239647786.

[232] Li, L.; Shih, C.; Yang, J.; Wang, L.; Li, D.; Ren, D. (2021). "New amber record of Braconidae (Insecta: Hymenoptera) from the mid-Cretaceous of Myanmar". Cretaceous Research. 124: Article 104794. Bibcode:2021CrRes.12404794L. doi:10.1016/j.cretres.2021.104794. S2CID 233882169.

[Burmapyris-233] Jouault, C.; Perrichot, V.; Nel, A. (2021). "New flat wasps from mid-Cretaceous Burmese amber deposits highlight the bethylid antiquity and paleobiogeography (Hymenoptera: Chrysidoidea)" (PDF). Cretaceous Research. 123: Article 104772. Bibcode:2021CrRes.12304772J. doi:10.1016/j.cretres.2021.104772. S2CID 234041758.

[Burmoxyela-234] Zheng, Y.; Hu, H.; Zhang, H.; Chen, J.; Rasnitsyn, A. P.; Zhuo, D. (2021). "New genus and species of syspastoxyelid sawflies (Insecta, Hymenoptera) from the mid-Cretaceous Kachin amber with a review of the family Syspastoxyelidae". Cretaceous Research. 127: Article 104940. Bibcode:2021CrRes.12704940Z. doi:10.1016/j.cretres.2021.104940.

[235] Li, L.; Shih, C.; Yang, J.; Gao, Y.; Ren, D. (2021). "Two new wasps from mid-Cretaceous Myanmar amber (Hymenoptera: Apocrita)". Cretaceous Research. 131: Article 105115. doi:10.1016/j.cretres.2021.105115. S2CID 244885445.

[Buserphites-236] Herbert, M. C. M.; McKellar, R. C. (2021). "New genera Buserphites and Mesoserphites (Hymenoptera: Serphitidae) from mid-Cretaceous amber of Myanmar" (PDF). Cretaceous Research. 130: Article 105025. doi:10.1016/j.cretres.2021.105025. S2CID 240331883.

[Chainochora-237] Chen, H.; van Achterberg, C.; Hong, P. (2021). "Two new genera of Protobraconinae (Hymenoptera, Braconidae) from mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 126: Article 104914. Bibcode:2021CrRes.12604914C. doi:10.1016/j.cretres.2021.104914.

[Gloxinius-238] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m Colombo, W. D.; Perkovsky, E. E.; Waichert, C.; Azevedo, C. O. (2021). "Synopsis of the fossil flat wasps Epyrinae (Hymenoptera, Bethylidae), with description of three new genera and 10 new species". Journal of Systematic Palaeontology. 19 (1): 39–89. doi:10.1080/14772019.2021.1882593. S2CID 232313171.

[Christophus-239] Brazidec, M.; Perrichot, V.; Nel, A.; Jouault, C. (2021). "Two new genera of belytine wasps (Hymenoptera: Diapriidae) from Cretaceous Burmese amber" (PDF). Proceedings of the Geologists' Association. 132 (5): 597–604. Bibcode:2021PrGA..132..597B. doi:10.1016/j.pgeola.2021.09.006. S2CID 244613026.

[240] Jouault, C.; Perrichot, V.; Nel, A. (2021). "Taxonomic addition to the early diverged parasitic wasps (Hymenoptera: Orussoidea) from the mid-Cretaceous Burmese amber" (PDF). Proceedings of the Geologists' Association. 132 (3): 324–331. Bibcode:2021PrGA..132..324J. doi:10.1016/j.pgeola.2021.02.002. S2CID 234848669.

[241] Jouault, C.; Rasnitsyn, A. P.; Pouillon, J.-M.; Nel, A. (2021). "The first Curiosivespa Rasnitsyn, 1975 (Hymenoptera: Vespidae) from the Lower Cretaceous Crato formation (Brazil)". Annales de la Société Entomologique de France. Nouvelle Série. 57 (6): 509–513. doi:10.1080/00379271.2021.1976672. S2CID 244229524.

[242] Santer, M.; Álvarez-Parra, S.; Nel, A.; Peñalver, E.; Delclòs, X. (2021). "New insights into the enigmatic Cretaceous family Spathiopterygidae (Hymenoptera: Diaprioidea)". Cretaceous Research. 133: Article 105128. doi:10.1016/j.cretres.2021.105128. hdl:2445/182542. S2CID 245556639.

[243] Radchenko, A. G. (2021). "New species of the fossil ant genus Drymomyrmex (Hymenoptera, Formicidae, Formicinae) from the late Eocene Rovno amber (Ukraine)". Palaeoentomology. 4 (6): 544–549. doi:10.11646/palaeoentomology.4.6.4. S2CID 245149637.

[244] Belokobylskij, S. A.; Dubovikoff, D. A.; Manukyan, A. R.; Zharkov, D. M. (2021). "Braconid parasitoids of ants (Hymenoptera, Braconidae, Euphorinae, Neoneurini) from Baltic amber with a discussion of records of fossil larvae parasitizing ant workers". Journal of Hymenoptera Research. 84: 29–43. doi:10.3897/jhr.84.67749.

[CRLietalAulacidae-245] Li, L.; Shih, P. J. M.; Zhang, X.; Peng, C.; Li, D.; Ren, D. (2021). "Geometric Morphometric Analysis of two genera confirm three new wasps from the mid-Cretaceous of Myanmar (Hymenoptera: Aulacidae)". Cretaceous Research. 130: Article 105057. doi:10.1016/j.cretres.2021.105057. S2CID 242552161.

[Palaeoentomology41Embolemus-246] Olmi, M.; Guglielmino, A.; Müller, P.; Capradossi, L.; Perkovsky, E. E. (2021). "Descriptions of the first females of Embolemus (Hymenoptera: Chrysidoidea) from Kachin (Myanmar) amber". Palaeoentomology. 4 (1): 44–49. doi:10.11646/palaeoentomology.4.1.9. S2CID 233944242.

[247] Jouault, C. (2021). "Mid-Cretaceous Burmese amber pelecinid wasps (Hymenoptera, Pelecinidae) support the hypothesis of an Asian origin of the family" (PDF). Annales de Paléontologie. 107 (1): Article 102464. Bibcode:2021AnPal.10702464J. doi:10.1016/j.annpal.2020.102464. S2CID 234321579.

[248] Archibald, S. B.; Aase, A.; Nel, A. (2021). "The second North American fossil horntail wood-wasp (Hymenoptera: Siricidae), from the early Eocene Green River Formation". Zootaxa. 4999 (4): 325–334. doi:10.11646/zootaxa.4999.4.2. PMID 34810481. S2CID 237762599.

[249] Davidian, E. M.; Kaliuzhna, M. O.; Perkovsky, E. E. (2021). "First aphidiine wasp from the Sakhalinian amber". Acta Palaeontologica Polonica. 66 (Supplement to 3): S59–S65. doi:10.4202/app.00843.2020. S2CID 236723591.

[250] Radchenko, A. G.; Perkovsky, E. E.; Vasilenko, D. V. (2021). "Formica species (Hymenoptera, Formicidae, Formicinae) in late Eocene Rovno amber". Journal of Hymenoptera Research. 82: 237–251. doi:10.3897/jhr.82.64599.

[251] Zheng, Y.; Kopylov, D. S.; Zhang, Q.; Zhang, H.-G.; Qin, M.; Chen, J.; Zhang, H.-C. (2021). "Discovery of a new symphytan wasp in the mid-Cretaceous Kachin amber (Hymenoptera, Syspastoxyelidae)". Palaeoentomology. 4 (6): 629–636. doi:10.11646/palaeoentomology.4.6.12. S2CID 245173388.

[Heteroichneumon-252] Kopylov, D. S.; Zhang, Q.; Zhang, H.-C. (2021). "The Darwin wasps (Hymenoptera: Ichneumonidae) of Burmese amber". Palaeoentomology. 4 (6): 592–603. doi:10.11646/palaeoentomology.4.6.8. S2CID 245157688.

[CRHolopsenella-253] Lepeco, A.; Melo, G. A. R. (2021). "The wasp genus †Holopsenella in mid-Cretaceous Burmese amber (Hymenoptera: †Holopsenellidae stat. nov.)". Cretaceous Research. 131: Article 105089. doi:10.1016/j.cretres.2021.105089. ISSN 0195-6671. S2CID 244055567.

[JHR81-254] Olmi, M.; Chen, H.-Y.; Shih, C.; Müller, P.; Capradossi, L.; Ren, D.; Perkovsky, E. E.; Guglielmino, A. (2021). "New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts". Journal of Hymenoptera Research. 81: 43–55. doi:10.3897/jhr.81.57792.

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[256] Bremer, J.; van de Kamp, T.; Talamas, E. J. (2021). "Janzenella theia Bremer & Talamas (Platygastroidea, Janzenellidae): a new species from Baltic amber". Journal of Hymenoptera Research. 87: 223–233. doi:10.3897/jhr.87.67256.

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[258] Jouault, C.; Maréchal, A.; Condamine, F. L.; Wang, B.; Nel, A.; Legendre, F.; Perrichot, V. (2022). "Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth–death process". Zoological Journal of the Linnean Society. 194 (4): 1396–1423. doi:10.1093/zoolinnean/zlab034.

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[260] Olmi, M.; Vasilenko, D. V.; Capradossi, L.; Perkovsky, E. E.; Guglielmino, A. (2021). "A new species of Lonchodryinus (Hymenoptera, Dryinidae) from Upper Eocene Baltic amber". Zootaxa. 5020 (2): 328–336. doi:10.11646/zootaxa.5020.2.5. PMID 34810999. S2CID 238658382.

[Megalyrhyssalus-261] Belokobylskij, S. A.; Jouault, C. (2021). "Two new striking braconid genera (Hymenoptera: Braconidae) from the mid-Cretaceous Burmese amber". Proceedings of the Geologists' Association. 132 (4): 426–437. Bibcode:2021PrGA..132..426B. doi:10.1016/j.pgeola.2021.04.003. S2CID 236271642.

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[266] Varela-Hernández, F.; Riquelme, F. (2021). "A New Ant Species of the Genus Pheidole Westwood, 1839 from Miocene Mexican Amber". Southwestern Entomologist. 46 (1): 75–82. doi:10.3958/059.046.0107. S2CID 232357684.

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[268] Jouault, C.; Nel, A. (2021). "A new species of Prosyntexis Sharkey, 1990 (Hymenoptera: Sepulcidae) from the Lower Cretaceous Crato Formation of Brazil confirmed by geometric morphometric analysis". Palaeoentomology. 4 (2): 171–177. doi:10.11646/palaeoentomology.4.2.6. S2CID 235511059.

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[Palaeoentomology46Rovenosa-272] Manukyan, A. R.; Zhindarev, L. A. (2021). "Fossil Darwin wasps (Hymenoptera: Ichneumonidae) from Baltic amber". Palaeoentomology. 4 (6): 637–647. doi:10.11646/palaeoentomology.4.6.13. S2CID 245191818.

[273] Simutnik, S. A.; Perkovsky, E. E.; Vasilenko, D. V. (2021). "Sakhalinencyrtus leleji Simutnik gen. et sp. nov. of earliest Encyrtidae (Hymenoptera, Chalcidoidea) from Sakhalinian amber". Journal of Hymenoptera Research. 84: 361–372. doi:10.3897/jhr.84.66367.

[274] Davidian, E. M.; Manukyan, A. R.; Belokobylskij, S. A. (2021). "A new genus and species of the aphid parasitoids (Hymenoptera: Braconidae: Aphidiinae) from the Eocene Sakhalinian amber". Palaeoentomology. 4 (6): 537–543. doi:10.11646/palaeoentomology.4.6.3. S2CID 245166207.

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[280] Zhuang, Y.; Ran, H.; Li, X.; Feng, C.; Liu, Y. (2021). "A new species of the iron maiden ant based on an alate female from mid-Cretaceous Burmese amber (Hymenoptera: Formicidae: †Zigrasimecia)". Cretaceous Research. 130: Article 105056. doi:10.1016/j.cretres.2021.105056. S2CID 240812443.

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[306] Godunko, R. J.; Martynov, A. V.; Staniczek, A. H. (2021). "First fossil record of the mayfly family Vietnamellidae (Insecta, Ephemeroptera) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution". ZooKeys (1036): 99–120. doi:10.3897/zookeys.1036.66435. PMC 8128840. PMID 34040482.

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[NJGPA3003-403] Nel, A.; Poschmann, M. J. (2021). "Early Permian acercarian insects (Miomoptera, Palaeomanteidae) from the Saar-Nahe Basin (Rhineland-Palatinate, SW Germany)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 300 (3): 267–277. doi:10.1127/njgpa/2021/0989. S2CID 236233140.

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[405] Falières, E.; Engel, M. S.; Nel, A. (2021). "Earliest occurrence of Embiidae: A new genus from earliest Eocene Oise amber (Insecta: Embiodea)". Comptes Rendus Palevol. 20 (38): 799–805. doi:10.5852/cr-palevol2021v20a38.

[Multivena-406] Lai, Z.; Yang, Q.; Shi, C.; Zhang, W.; Ren, D. (2021). "New webspinners (Insecta: Embioptera) from Upper Cretaceous amber of northern Myanmar". Cretaceous Research. 130: Article 105040. doi:10.1016/j.cretres.2021.105040. S2CID 239202878.

[Kamopanorpa-407] Bashkuev, A. S.; Sukatsheva, I. D. (2021). "New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)". Palaeoentomology. 4 (5): 453–461. doi:10.11646/palaeoentomology.4.5.11. ISSN 2624-2826. S2CID 240525412.

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