Harpalus (beetle)
| Harpalus | |
|---|---|
| |
| Harpalus affinis | |
Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Coleoptera |
| Family: | Carabidae |
| Subfamily: | Harpalinae |
| Tribe: | Harpalini |
| Genus: | Harpalus Latreille, 1802 |
| Synonyms | |
| |
Harpalus is a genus of ground beetle first described by Pierre André Latreille in 1802.[1]
Species
Harpalus contains the following 430 species:[2][3]
- Subgenus Cephalophonus Ganglbauer, 1891
- Harpalus cephalotes Fairmaire & Laboulbène, 1854
- Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982
- Harpalus agnatus Reiche, 1850
- Harpalus cyrenaicus Koch, 1939
- Harpalus fulvus Dejean, 1829
- Harpalus grilli Kataev, 2002
- Harpalus idiotus Bates, 1889
- Harpalus janinae Jeanne, 1984
- Harpalus litigiosus Dejean, 1829
- Harpalus melancholicus Dejean, 1829
- Harpalus schaumii Wollaston, 1864
- Harpalus tenebrosus Dejean, 1829
- Subgenus Glanodes Casey, 1914
- Harpalus cohni Ball, 1972
- Harpalus corpulentus (Casey, 1914)
- Harpalus huachuca Ball, 1972
- Harpalus obliquus G.Horn, 1880
- Harpalus puncticeps (Casey, 1914)
- Harpalus stephani Ball, 1972
- Subgenus Harpalus Latreille, 1802
- Harpalus acupalpoides Reitter, 1900
- Harpalus aeneipennis (Faldermann, 1836)
- Harpalus aequicollis Motschulsky, 1844
- Harpalus affinis (Schrank, 1781)
- Harpalus agakhaniantzi (Mikhailov, 1972)
- Harpalus akbensis Jedlicka, 1958
- Harpalus akinini Tschitscherine, 1895
- Harpalus alajensis Tschitscherine, 1898
- Harpalus albanicus Reitter, 1900
- Harpalus alexandrae Kataev, 1990
- Harpalus alexeevi Kataev, 1990
- Harpalus alpivagus Tschitscherine, 1899
- Harpalus amarellus Bates, 1891
- Harpalus amariformis Motschulsky, 1844
- Harpalus amplicollis Ménétriés, 1848
- Harpalus amputatus Say, 1830
- Harpalus anatolicus Tschitscherine, 1898
- Harpalus angulatus Putzeys, 1878
- Harpalus angustipennis Boheman, 1848
- Harpalus angustitarsis Reitter, 1887
- Harpalus animosus Casey, 1924
- Harpalus anisodactyliformis Solsky, 1874
- Harpalus antonowi (Tschitscherine, 1898)
- Harpalus anxioides Kataev, 1991
- Harpalus anxius (Duftschmid, 1812)
- Harpalus araraticus Mlynar, 1979
- Harpalus arcuatus Tschitscherine, 1898
- Harpalus arnoldii Kataev, 1988
- Harpalus ascetes Kataev & Wrase, 1997
- Harpalus asemus Basilewsky, 1946
- Harpalus aterrimus Casey, 1914
- Harpalus atratus Latreille, 1804
- Harpalus atrichatus Hatch, 1949
- Harpalus atripes Casey, 1914
- Harpalus attenuatus Stephens, 1828
- Harpalus australasiae Dejean, 1829
- Harpalus autumnalis (Duftschmid, 1812)
- Harpalus badakschanus Jedlicka, 1956
- Harpalus baleensis Clarke, 1973
- Harpalus balli Noonan, 1991
- Harpalus basanicus J.Sahlberg, 1913
- Harpalus basilewskyi Facchini, 2003
- Harpalus basuto Basilewsky, 1958
- Harpalus bellieri Reiche, 1861
- Harpalus beneshi Kataev & Wrase, 1997
- Harpalus brachypterus Tschitscherine, 1898
- Harpalus brevicornis Germar, 1823
- Harpalus brevis Motschulsky, 1844
- Harpalus breviusculus Chaudoir, 1846
- Harpalus bucharicus Tschitscherine, 1898
- Harpalus bulirschi (Facchini, 2012)
- Harpalus bungii Chaudoir, 1844
- Harpalus caeruleatus Bates, 1878
- Harpalus caiphus Reiche & Saulcy, 1855
- Harpalus calathoides Motschulsky, 1844
- Harpalus capicola Dejean, 1829
- Harpalus cardoni Antoine, 1922
- Harpalus caspius (Steven, 1806)
- Harpalus cautus Dejean, 1829
- Harpalus chalcentus Bates, 1873
- Harpalus chasanensis Lafer, 1989
- Harpalus chobautianus Lutshnik, 1922
- Harpalus chrysopus Reitter, 1887
- Harpalus circumpunctatus Chaudoir, 1846
- Harpalus cisteloides Motschulsky, 1844
- Harpalus clarkei Kataev & J.Schmidt, 2020
- Harpalus compressus Motschulsky, 1844
- Harpalus contemptus Dejean, 1829
- Harpalus convexus Faldermann, 1836
- Harpalus corporosus (Motschulsky, 1862)
- Harpalus corrugatus Basilewsky, 1958
- Harpalus crates Bates, 1883
- Harpalus cupreus Dejean, 1829
- Harpalus cursorius Péringuey, 1896
- Harpalus cyanopterus (Tschitscherine, 1897)
- Harpalus cyclogonus Chaudoir, 1844
- Harpalus danieli Reitter, 1900
- Harpalus davidianus Tschitscherine in Jakobson, 1903
- Harpalus decipiens Dejean, 1829
- Harpalus defector Péringuey, 1896
- Harpalus desertus LeConte, 1859
- Harpalus diligens Tschitscherine, 1898
- Harpalus dimidiatus (P.Rossi, 1790)
- Harpalus dispar Dejean, 1829
- Harpalus dissitus Antoine, 1931
- Harpalus distinguendus (Duftschmid, 1812)
- Harpalus diversicollis Basilewsky, 1958
- Harpalus dubius Boheman, 1848
- Harpalus dudkoi Kataev, 2011
- Harpalus durangoensis Bates, 1891
- Harpalus ebeninus Heyden, 1870
- Harpalus egenus Dejean, 1829
- Harpalus egorovi Lafer, 1989
- Harpalus ellipsis LeConte, 1847
- Harpalus elliptipennis Facchini, 2003
- Harpalus erosus Mannerheim, 1825
- Harpalus exiguus Boheman, 1848
- Harpalus famelicus Tschitscherine, 1898
- Harpalus farkaci Kataev & Wrase, 1995
- Harpalus fimetarius Dejean, 1829
- Harpalus flavescens (Piller & Mitterpacher, 1783)
- Harpalus flavicornis Dejean, 1829
- Harpalus foveiger Tschitscherine, 1895
- Harpalus franzi Mateu, 1954
- Harpalus frater Chaudoir, 1876
- Harpalus fraternus LeConte, 1852
- Harpalus froelichii Sturm, 1818
- Harpalus fulvicornis Thunberg, 1806
- Harpalus fulvilabris Mannerheim, 1853
- Harpalus fulvipennis Chaudoir, 1843
- Harpalus fuscicornis Ménétriés, 1832
- Harpalus fuscipalpis Sturm, 1818
- Harpalus fuscipennis Wiedemann, 1825
- Harpalus fuscoaeneus Dejean, 1829
- Harpalus ganssuensis Semenov, 1889
- Harpalus giacomazzoi Kataev & Wrase, 1996
- Harpalus gilgil Basilewsky, 1946
- Harpalus gisellae Csiki, 1932
- Harpalus glasunovi Kataev, 1987
- Harpalus gravis LeConte, 1858
- Harpalus gregoryi Alluaud, 1917
- Harpalus hartmanni Kataev, 2002
- Harpalus herbivagus Say, 1823
- Harpalus heyrovskyi Jedlicka, 1928
- Harpalus hiekei Kataev & Wrase, 2010
- Harpalus hirtipes (Panzer, 1796)
- Harpalus honestus (Duftschmid, 1812)
- Harpalus hospes Sturm, 1818
- Harpalus hybridus Boheman, 1848
- Harpalus impressus Roth, 1851
- Harpalus inconcinnus Chaudoir, 1876
- Harpalus indianus Csiki, 1932
- Harpalus indicola Bates, 1878
- Harpalus indigens Casey, 1924
- Harpalus inexspectatus Kataev, 1989
- Harpalus ingenuus Tschitscherine, 1898
- Harpalus innocuus LeConte, 1863
- Harpalus italus Schaum, 1860
- Harpalus jeanneli Basilewsky, 1946
- Harpalus jordanus Jedlicka, 1964
- Harpalus kabakianus Kataev, 1988
- Harpalus kabakovi Kataev, 1987
- Harpalus kadleci Kataev & Wrase, 1995
- Harpalus kadyrbekovi Kataev, 1988
- Harpalus kagyzmanicus Kataev, 1984
- Harpalus kandaharensis Jedlicka, 1956
- Harpalus karakorum Jedlicka, 1958
- Harpalus karamani Apfelbeck, 1902
- Harpalus kazanensis Jedlicka, 1958
- Harpalus kaznakovi Kataev & Wrase, 1997
- Harpalus kibonoti Alluaud, 1927
- Harpalus kirgisicus (Motschulsky, 1844)
- Harpalus kiritshenkoi Kataev, 1990
- Harpalus klapperichi Jedlicka, 1956
- Harpalus kmecoi Facchini, 2003
- Harpalus kozlovi Kataev, 1993
- Harpalus kryzhanovskii Kataev, 1988
- Harpalus kunarensis Kataev, 1993
- Harpalus laetus Reiche, 1843
- Harpalus laevipes Zetterstedt, 1828
- Harpalus lama Kataev & Wrase, 1997
- Harpalus lateralis Dejean, 1829
- Harpalus laticeps LeConte, 1850
- Harpalus latus (Linnaeus, 1758)
- Harpalus lederi Tschitscherine, 1899
- Harpalus lethierryi Reiche, 1860
- Harpalus lewisii LeConte, 1865
- Harpalus longipalmatus Mordkovitsh, 1969
- Harpalus lopezi J.Serrano & Lencina, 2009
- Harpalus lugubris Boheman, 1848
- Harpalus lumbaris Mannerheim, 1825
- Harpalus luteicornis (Duftschmid, 1812)
- Harpalus lutshniki Schauberger, 1932
- Harpalus macronotus Tschitscherine, 1893
- Harpalus mairei Peyerimhoff, 1928
- Harpalus major (Motschulsky, 1850)
- Harpalus makhekensis Basilewsky, 1958
- Harpalus manas Kataev, 1990
- Harpalus marginellus Gyllenhal, 1827
- Harpalus martini Van Dyke, 1926
- Harpalus masoreoides Bates, 1878
- Harpalus massarti Burgeon, 1936
- Harpalus mauritanicus Gaubil, 1844
- Harpalus medvedevi Kataev, 2006
- Harpalus megacephalus LeConte, 1847
- Harpalus melaneus Bates, 1878
- Harpalus metallinus Ménétriés, 1836
- Harpalus metarsius Andrewes, 1930
- Harpalus meteorus Basilewsky, 1946
- Harpalus michaili Kataev, 1990
- Harpalus michailovi Kataev, 1987
- Harpalus microthorax (Motschulsky, 1849)
- Harpalus miles Péringuey, 1896
- Harpalus minutissimus Facchini, 2015
- Harpalus minutulus Kataev & Liang, 2004
- Harpalus mitridati Pliginskiy, 1915
- Harpalus mlynari Kataev, 1990
- Harpalus modestus Dejean, 1829
- Harpalus morvani Kataev, 2002
- Harpalus murzini Kataev, 2008
- Harpalus nanniscus Péringuey, 1896
- Harpalus natalensis Boheman, 1848
- Harpalus natalicus Péringuey, 1896
- Harpalus neglectus Audinet-Serville, 1821
- Harpalus nevadensis K. & J.Daniel, 1898
- Harpalus nigrans A.Morawitz, 1862
- Harpalus nigritarsis C.R.Sahlberg, 1827
- Harpalus numidicus Bedel, 1893
- Harpalus nyassicus Basilewsky, 1946
- Harpalus oblitus Dejean, 1829
- Harpalus obnixus Casey, 1924
- Harpalus ochropus Kirby, 1837
- Harpalus oodioides Dejean, 1829
- Harpalus opacipennis (Haldeman, 1843)
- Harpalus optabilis Dejean, 1829
- Harpalus ovtshinnikovi Kataev, 1990
- Harpalus pallidipennis A.Morawitz, 1862
- Harpalus pallipes Chaudoir, 1837
- Harpalus parallelocollis Facchini, 2003
- Harpalus parasinuatus Kataev & Liang, 2007
- Harpalus parvulus Dejean, 1829
- Harpalus petri Tschitscherine, 1902
- Harpalus picipennis (Duftschmid, 1812)
- Harpalus plancyi Tschitscherine, 1897
- Harpalus plenalis Casey, 1914
- Harpalus politus Dejean, 1829
- Harpalus polyglyptus Schaum, 1862
- Harpalus potanini Tschitscherine, 1906
- Harpalus praecurrens Schauberger, 1934
- Harpalus procognatus Lorenz, 1998
- Harpalus progrediens Schauberger, 1922
- Harpalus prosperus Basilewsky, 1972
- Harpalus providens Casey, 1914
- Harpalus pseudoserripes Reitter, 1900
- Harpalus pterostichus (Reitter, 1900)
- Harpalus puetzi Kataev & Wrase, 1997
- Harpalus pulchrinulus Reitter, 1900
- Harpalus pulvinatus Ménétriés, 1848
- Harpalus pumilus Sturm, 1818
- Harpalus punctatostriatus Dejean, 1829
- Harpalus punctipennis Mulsant, 1852
- Harpalus pusillus (Motschulsky, 1850)
- Harpalus pygmaeus Dejean, 1829
- Harpalus quadratus Chaudoir, 1846
- Harpalus raphaili Kataev, 1997
- Harpalus reflexus Putzeys in Schneider & Leder, 1878
- Harpalus reitteri Wrase & Kataev, 2011
- Harpalus remboides Solsky, 1874
- Harpalus reversus Casey, 1924
- Harpalus rhemboides Solsky, 1874
- Harpalus rivalsi Jeannel, 1948
- Harpalus rotundus Facchini, 2003
- Harpalus rougemonti Clarke, 1973
- Harpalus rubripes (Duftschmid, 1812)
- Harpalus rufipalpis Sturm, 1818
- Harpalus rufiscapus Gebler, 1833
- Harpalus rufitarsoides Schauberger, 1934
- Harpalus rufomarginatus (Boheman, 1848)
- Harpalus rumelicus Apfelbeck, 1904
- Harpalus salinulus Reitter, 1900
- Harpalus salinus Dejean, 1829
- Harpalus sanctaehelenae Basilewsky, 1972
- Harpalus saxicola Dejean, 1829
- Harpalus semenowi Tschitscherine, 1901
- Harpalus semipunctatus Dejean, 1829
- Harpalus serripes (Quensel, 1806)
- Harpalus servus (Duftschmid, 1812)
- Harpalus setiporus (Reitter, 1894)
- Harpalus siculus Dejean, 1829
- Harpalus silipes Dejean, 1831
- Harpalus sinuaticollis Facchini, 2015
- Harpalus sinuatus Tschitscherine, 1893
- Harpalus smaragdinus (Duftschmid, 1812)
- Harpalus smyrnensis Heyden, 1888
- Harpalus solitaris Dejean, 1829
- Harpalus somereni Basilewsky, 1946
- Harpalus somnulentus Dejean, 1829
- Harpalus spadiceus Dejean, 1829
- Harpalus spretus Péringuey, 1896
- Harpalus spurius Péringuey, 1896
- Harpalus stevenii Dejean, 1829
- Harpalus stevensi Kataev, 2011
- Harpalus stoetznerianus Schauberger, 1932
- Harpalus strenuus Tschitscherine, 1898
- Harpalus subaeneus Boheman, 1848
- Harpalus subcylindricus Dejean, 1829
- Harpalus subtruncatus Chaudoir, 1846
- Harpalus sulphuripes Germar, 1823
- Harpalus sundaicus Schauberger, 1933
- Harpalus sushenicus Kataev, 1990
- Harpalus suturangulus Reitter, 1887
- Harpalus szalliesi Kataev & Wrase, 1995
- Harpalus taciturnus Dejean, 1829
- Harpalus tangutorum Kataev, 1993
- Harpalus tardus (Panzer, 1796)
- Harpalus tarsalis Mannerheim, 1825
- Harpalus terrestris (Motschulsky, 1844)
- Harpalus tibeticus Andrewes, 1930
- Harpalus tichonis Jakobson, 1907
- Harpalus tiridates Reitter, 1900
- Harpalus tithonus Reitter, 1900
- Harpalus tjanschanicus Semenov, 1889
- Harpalus torosensis Jedlicka, 1961
- Harpalus torridoides Reitter, 1900
- Harpalus trichophorus Tschitscherine, 1897
- Harpalus triseriatus A.Fleischer, 1897
- Harpalus turcicus Jedlicka, 1958
- Harpalus turmalinus Erichson, 1847
- Harpalus udege Lafer, 1989
- Harpalus uniformis (Motschulsky, 1844)
- Harpalus ussuricus Mlynar, 1979
- Harpalus vanemdeni Schauberger, 1932
- Harpalus venator Boheman, 1848
- Harpalus ventralis LeConte, 1847
- Harpalus vereschaginae Kataev, 1988
- Harpalus vernicosus Kataev & Liang, 2007
- Harpalus viridanus (Motschulsky, 1844)
- Harpalus viridicupreus Reiche, 1843
- Harpalus vittatus Gebler, 1833
- Harpalus wadiensis Jedlicka, 1964
- Harpalus wohlberedti Emden, 1932
- Harpalus xanthopus Gemminger & Harold, 1868
- Harpalus zabroides Dejean, 1829
- Harpalus zhdankoi Kataev, 1990
- Subgenus Loboharpalus Schauberger, 1932
- Harpalus platynotus Bates, 1873
- Harpalus rubefactus Bates, 1873
- Subgenus Opadius Casey, 1914
- Harpalus apache Kataev, 2010
- Harpalus cordatus (LeConte, 1853)
- Harpalus cordifer Notman, 1919
- Subgenus Plectralidus Casey, 1914
- Harpalus erraticus Say, 1823
- Harpalus retractus LeConte, 1863
- Subgenus Pseudoophonus Motschulsky, 1844
- Harpalus actiosus Casey, 1914
- Harpalus aenigma (Tschitscherine, 1897)
- Harpalus alienus Bates, 1878
- Harpalus aogashimensis (Habu, 1957)
- Harpalus azumai Habu, 1968
- Harpalus babai Habu, 1973
- Harpalus calceatus (Duftschmid, 1812)
- Harpalus caliginosus (Fabricius, 1775)
- Harpalus capito A.Morawitz, 1862
- Harpalus compar LeConte, 1847
- Harpalus coreanus (Tschitscherine, 1895)
- Harpalus davidi (Tschitscherine, 1897)
- Harpalus disimuciulus Huang; Lei; Yan & Hu, 1996
- Harpalus eous Tschitscherine, 1901
- Harpalus erythropus Dejean, 1829
- Harpalus faunus Say, 1823
- Harpalus fokienensis Schauberger, 1930
- Harpalus griseus (Panzer, 1796)
- Harpalus hatchi Ball & Anderson, 1962
- Harpalus hauserianus Schauberger, 1929
- Harpalus indicus Bates, 1891
- Harpalus jureceki (Jedlicka, 1928)
- Harpalus katiae F.Battoni, 1985
- Harpalus liobasis Chaudoir, 1868
- Harpalus meghalayensis Kataev, 2001
- Harpalus meridianus Andrewes, 1923
- Harpalus paratus Casey, 1924
- Harpalus pastor Motschulsky, 1844
- Harpalus pensylvanicus (DeGeer, 1774)
- Harpalus poncei Will, 2002
- Harpalus protractus Casey, 1914
- Harpalus pseudohauserianus Kataev, 2001
- Harpalus pseudophonoides Schauberger, 1930
- Harpalus roninus Bates, 1873
- Harpalus rufipes (DeGeer, 1774)
- Harpalus sericatus Tschitscherine, 1906
- Harpalus simplicidens Schauberger, 1929
- Harpalus singularis Tschitscherine, 1906
- Harpalus sinicus Hope, 1845
- Harpalus suensoni Kataev, 1997
- Harpalus texanus Casey, 1914
- Harpalus tridens A.Morawitz, 1862
- Harpalus ussuriensis Chaudoir, 1863
- Harpalus vagans LeConte, 1865
- Harpalus yinchuanensis Huang, 1993
- Subgenus Semiophonus Schauberger, 1933
- Harpalus signaticornis (Duftschmid, 1812)
- Subgenus Zangoharpalus Huang, 1998
- Harpalus mariae Kataev, 1997
- Harpalus microdemas Schauberger, 1932
- Harpalus praticola Bates, 1891
- Harpalus pseudotinctulus Schauberger, 1932
- Harpalus tinctulus Bates, 1873
- Extinct, not assigned to subgenus
- †Harpalus abolitus C. & L.Heyden, 1866
- †Harpalus anactus Giebel, 1856
- †Harpalus bruckmanni Heer, 1862
- †Harpalus burmeisteri Giebel, 1856
- †Harpalus conditus Scudder, 1900
- †Harpalus constrictus Heer, 1862
- †Harpalus deletus Oustalet, 1864
- †Harpalus dilatatus Theobald, 1937
- †Harpalus ewaldi Giebel, 1856
- †Harpalus excavatus Forster, 1891
- †Harpalus heeri Giebel, 1856
- †Harpalus knorri Giebel, 1856
- †Harpalus lheritieri Piton, 1940
- †Harpalus liasinus Giebel, 1856
- †Harpalus maceratus Wickham, 1911
- †Harpalus nero Oustalet, 1864
- †Harpalus nitens Birket-Smith, 1977
- †Harpalus nuperus Scudder, 1900
- †Harpalus obtusus Forster, 1891
- †Harpalus oustaleti Meunier, 1915
- †Harpalus pleistocenicus Lomnicki, 1894
- †Harpalus redivivus Wickham, 1917
- †Harpalus schlotheimi Giebel, 1856
- †Harpalus sinis Heer, 1847
- †Harpalus stierlini Heer, 1862
- †Harpalus stygius Heer, 1862
- †Harpalus tabidus Heer, 1847
- †Harpalus tardigradus Heer, 1862
- †Harpalus ulomaeformis Wickham, 1917
- †Harpalus veterum Cockerell, 1920
- †Harpalus whitfieldi Scudder, 1900
References
- ^ Latreille, Pierre André (1802). "Histoire naturelle, générale et particulière des crustacés et des insectes". 3. F. Dufart. doi:10.5962/bhl.title.15764. OCLC 5050710.
{{cite journal}}: Cite journal requires|journal=(help) - ^ "Harpalus Latreille, 1802". Catalogue of Life. Retrieved 2023-04-09.
- ^ Lorenz, Wolfgang (2021). "Carabcat Database". doi:10.48580/dfqf-3dk. Retrieved 2023-03-04.